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Taxon	Taxonomy ID	Rank	Common Name
Scarabaeoidea ↑	75546	superfamily
Belohinidae	614208	family
Belohina	3085238	genus
- Belohina inexpectata	3085239	species
Bolboceratidae	110049	family
Bolboceras	463063	genus
unclassified Bolboceras	2624483	no rank
- Bolboceras sp.	2999166	species
- Bolboceras sp. 1 YvdM-2007	463067	species
- Bolboceras sp. 2 YvdM-2007	463068	species
Elephastomus	295528	genus
- Elephastomus proboscideus	295529	species
unclassified Elephastomus	2642373	no rank
- Elephastomus sp. NLG-2014	1551888	species
Eucanthus	463065	genus
- Eucanthus impressus	2680422	species
- Eucanthus lazarus	558330	species
Odonteus	396634	genus
- Odonteus armiger	1542177	species
- Odonteus liebecki	1770658	species
- Odonteus thoracicornis	2680752	species
unclassified Odonteus	2622312	no rank
- Odonteus sp. BMNH 677839	1142413	species
Prototrupes	1142445	genus
unclassified Prototrupes	2645899	no rank
- Prototrupes sp. BMNH 793036	1142446	species
unclassified Bolboceratidae	1461477	no rank
- Bolboceratidae sp. B1	1461481	species
- Bolboceratidae sp. B2	1461480	species
- Bolboceratidae sp. B3	1461479	species
- Bolboceratidae sp. B4	1461478	species
Diphyllostomatidae	110047	family
Diphyllostoma	1542231	genus
- Diphyllostoma fimbriata	1542232	species
Geotrupidae	110052	family	earth-boring dung beetles
Anoplotrupes	260541	genus
- Anoplotrupes stercorosus	260542	species
unclassified Anoplotrupes	2908763	no rank
- Anoplotrupes sp.	2908764	species
Australobolbus	1551972	genus
unclassified Australobolbus	2634580	no rank
- Australobolbus sp. CSR137	2055595	species
- Australobolbus sp. FLSHS BIONAUT2017 Sample_6	2126063	species
- Australobolbus sp. NLG-2014	1551973	species
Blackbolbus	1551980	genus
unclassified Blackbolbus	2626058	no rank
- Blackbolbus sp. NLG-2014	1551981	species
Bolbocaffer	1142437	genus
unclassified Bolbocaffer	2620632	no rank
- Bolbocaffer sp. BMNH 793035	1142438	species
Bolboceratex	1205608	genus
unclassified Bolboceratex	2645599	no rank
- Bolboceratex sp. BOL01	1205609	species
Bolboceroides	1142439	genus
- Bolboceroides capensis	1142440	species
Bolbocerosoma	2680313	genus
- Bolbocerosoma confusum	2680314	species
Bolborhinum	1627857	genus
- Bolborhinum geotrupoides	1627858	species
Bolbotrypes	1290872	genus
- Bolbotrypes davidis	1290873	species
Ceratophyus	997837	genus
- Ceratophyus hoffmannseggi	997838	species
- Ceratophyus martinezi	997839	species
- Ceratophyus polyceros	1627842	species
Frickius	351536	genus
- Frickius variolosus	997831	species
Geotrupes	7086	genus
- Geotrupes balyi	2109416	species
- Geotrupes blackburnii	3291020	species
- Geotrupes hornii	2109417	species
- Geotrupes ibericus	997832	species
- Geotrupes mutator	166344	species
- Geotrupes opacus	3082343	species
- Geotrupes semiopacus	2680486	species
- Geotrupes spiniger	295525	species
- Geotrupes stercorarius	166345	species
unclassified Geotrupes	2622733	no rank
- Geotrupes sp. DDM_3308	2652989	species
- Geotrupes sp. INB097	2055414	species
- Geotrupes sp. NLG-2014	1551893	species
Gilletinus	1552014	genus
unclassified Gilletinus	2618724	no rank
- Gilletinus sp. NLG-2014	1552015	species
Haplogeotrupes	997840	genus
- Haplogeotrupes guatemalensis	997841	species
Jekelius	997842	genus
- Jekelius albarracinus	997843	species
- Jekelius balearicus	997844	species
- Jekelius brullei	997845	species
- Jekelius castillanus	997846	species
- Jekelius catalonicus	997847	species
- Jekelius hernandezi	997850	species
- Jekelius hispanus	997851	species
- Jekelius intermedius	997852	species
- Jekelius nitidus	997855	species
- Jekelius punctatolineatus	997856	species
- Jekelius punctulatus	997857	species
Lethrus	295530	genus
- Lethrus anisodon	2609320	species
- Lethrus apterus	1612427	species
- Lethrus ares	2609321	species
- Lethrus bituberculatus	2609322	species
- Lethrus elephas	1844510	species
- Lethrus geminatus	1542163	species
- Lethrus halkidikiensis	2609323	species
- Lethrus inermis	2609324	species
- Lethrus karelini	295531	species
- Lethrus kentauensis	2609325	species
- Lethrus perun	1844511	species
- Lethrus raymondi	1004725	species
- Lethrus schaumi	2609326	species
- Lethrus schneideri	2609327	species
- Lethrus scoparius	2609328	species
- Lethrus strymonensis	2609329	species
- Lethrus superbus	2609330	species
Odontaeus	465314	genus
unclassified Odontaeus	2638435	no rank
- Odontaeus sp. BMNH 677839	465315	species
Onthotrupes	997860	genus
- Onthotrupes herbeus	997861	species
- Onthotrupes nebularum	997862	species
Phelotrupes	452575	genus
- Phelotrupes auratus	762646	species
- Phelotrupes formosanus	1689729	species
- Phelotrupes laevistriatus	273926	species
- Phelotrupes oberthuri	3128455	species
- Phelotrupes oshimanus	452577	species
Sericotrupes	997863	genus
- Sericotrupes niger	997864	species
Silphotrupes	997865	genus
- Silphotrupes escorialensis	1844509	species
- Silphotrupes orocantabricus	997866	species
- Silphotrupes punctatissimus	997867	species
Taurocerastes	997868	genus
- Taurocerastes patagonicus	997869	species
Thorectes	166373	genus
- Thorectes armifrons	997833	species
- Thorectes baraudi	997834	species
- Thorectes ferreri	997835	species
- Thorectes hoppei	1588511	species
- Thorectes laevigatus	3390091	species
- Thorectes lusitanicus	166374	species
- Thorectes sardous	3139941	species
- Thorectes trituberculatus	3390092	species
- Thorectes valencianus	997836	species
unclassified Thorectes	2999161	no rank
- Thorectes sp.	2999162	species
Trypocopris	997870	genus
- Trypocopris alpinus	1588524	species
- Trypocopris pyrenaeus	997871	species
- Trypocopris vernalis	1588525	species
unclassified Trypocopris	2634360	no rank
- Trypocopris sp. BMNH 793041	1142430	species
Typhaeus	166375	genus
- Typhaeus momus	166377	species
- Typhaeus typhoeoides	3390093	species
- Typhaeus typhoeus	166376	species	minotaur beetle
unclassified Geotrupidae	396586	no rank
- Bolbocerinae sp. BM677840	396587	species
Glaphyridae	110051	family
Amphicoma	2055586	genus
Amphicoma abdominalis	2722828	species
- Amphicoma abdominalis aemiliana	2722829	subspecies
- Amphicoma corinthia	2722826	species
- Amphicoma iberica	2722830	species
unclassified Amphicoma	2626977	no rank
- Amphicoma sp. INB011	2055587	species
Athypna	465180	genus
- Athypna carceli	465181	species
- Athypna meles	2722827	species
Eulasia	465235	genus
Eulasia arctos	2722831	species
- Eulasia arctos arctos	2722832	subspecies
- Eulasia arctos armeniaca	2722833	subspecies
- Eulasia bombylius	1661826	species
- Eulasia chalybaea	2722834	species
- Eulasia chrysopyga	2722835	species
- Eulasia daccordii	2722836	species
- Eulasia dilutipennis	2722837	species
- Eulasia fastuosa	2722838	species
- Eulasia genei	2722839	species
- Eulasia harmonia	2722840	species
- Eulasia korbi	2722841	species
- Eulasia nitidicollis	1142403	species
- Eulasia papaveris	1142404	species
- Eulasia pietschmanni	2722842	species
- Eulasia rapillyi	2722843	species
- Eulasia rittneri	2722844	species
- Eulasia saccai	2722845	species
Eulasia vittata	1142405	species
- Eulasia vittata lineata	2722846	subspecies
unclassified Eulasia	2677216	no rank
- Eulasia sp. BMNH 671327	465236	species
Glaphyrus	702049	genus
- Glaphyrus aulicus	1142407	species
- Glaphyrus caucasicus	2722847	species
- Glaphyrus comosus	1227469	species
- Glaphyrus opulentus	2722848	species
- Glaphyrus orbachi	2722849	species
- Glaphyrus serratulae	2722850	species
- Glaphyrus superbus	702050	species
Lichnanthe	295532	genus
- Lichnanthe apina	1542164	species
- Lichnanthe rathvoni	295533	species
Propomacrus	1142443	genus
- Propomacrus bimucronatus	1142444	species
- Propomacrus cypriacus	1968893	species
- Propomacrus davidi	1968894	species
Pygopleurus	1142447	genus
- Pygopleurus apicalis	2722860	species
- Pygopleurus besucheti	2722851	species
- Pygopleurus cf. syriacus	3425649	species
- Pygopleurus chrysonotus	2722853	species
- Pygopleurus despectus	2722852	species
- Pygopleurus foina	2802475	species
- Pygopleurus hirsutus	2722861	species
- Pygopleurus israelitus	2722854	species
- Pygopleurus kareli	2803042	species
- Pygopleurus katbehi	2722855	species
- Pygopleurus labaumei	2803043	species
- Pygopleurus libanonensis	2722856	species
- Pygopleurus medius	2803044	species
- Pygopleurus orientalis	1142448	species
- Pygopleurus ponticus	2722857	species
- Pygopleurus scutellatus	2722862	species
- Pygopleurus syriacus	2722858	species
- Pygopleurus transcaucasicus	2722859	species
- Pygopleurus vulpes	2803033	species
Glaresidae	110045	family
Glaresis	295534	genus
- Glaresis beckeri	1627846	species
- Glaresis ecostata	1542155	species
unclassified Glaresis	2644194	no rank
- Glaresis sp. AD-2015	1661494	species
- Glaresis sp. BMNH 793034	1142408	species
- Glaresis sp. BOLD:AAH0416	2659208	species
- Glaresis sp. GLA01	1205555	species
- Glaresis sp. TJH-2004	295535	species
unclassified Glaresidae	2917905	no rank
- Glaresidae sp. CNC COL EO119614	2917906	species
Hybosoridae	110053	family
Anaidinae	2606940	subfamily
Anaides	1142432	genus
- Anaides laticollis	2607238	species
unclassified Anaides	2627735	no rank
Chaetodus	2607016	genus
- Chaetodus asuai	2903156	species
- Chaetodus irregularis	2903159	species
- Chaetodus piceus	3085230	species
- Chaetodus teamscaraborum	3085231	species
Cryptogenius	2607017	genus
- Cryptogenius westwoodi	3085232	species
unclassified Cryptogenius	2607260	no rank
Ceratocanthinae	2606939	subfamily
Afrocloetus	2606941	genus
- Afrocloetus gibbosus	2606942	species
Aneilobolus	2606946	genus
unclassified Aneilobolus	2607254	no rank
Astaenomoechus	2606948	genus
- Astaenomoechus americanus	2953187	species
unclassified Astaenomoechus	2607256	no rank
Balleriodes	2943310	genus
- Balleriodes sphaera	2943311	species
Baloghianestes	2606950	genus
- Baloghianestes anceps	2607236	species
- Baloghianestes oribatidiformis	2607237	species
unclassified Baloghianestes	2607258	no rank
Carinophilharmostes	2606954	genus
- Carinophilharmostes vadoni	2607235	species
Ceratocanthus	1205612	genus
- Ceratocanthus amazonicus	2607239	species
- Ceratocanthus relucens	3085229	species
unclassified Ceratocanthus	2626782	no rank
Congomostes	2606943	genus
- Congomostes hintelmanni	2606944	species
Cryptophilharmostes	2606959	genus
- Cryptophilharmostes mahunkai	2607240	species
- Cryptophilharmostes merkli	2607241	species
unclassified Cryptophilharmostes	2607262	no rank
Cyphopisthes	1552001	genus
unclassified Cyphopisthes	2607265	no rank
Germarostes	1710472	genus
- Germarostes aphodioides	1710473	species
- Germarostes globosus	2953189	species
- Germarostes posticus	2903150	species
- Germarostes senegalensis	2903149	species
unclassified Germarostes	2607267	no rank
Ivieolus	2606965	genus
- Ivieolus brooksi	3085234	species
- Ivieolus inflaticollis	2953191	species
Madrasostes	2606966	genus
unclassified Madrasostes	2607269	no rank
Martinezostes	351538	genus
- Martinezostes fortecostatus	351539	species
- Martinezostes posticus	3085235	species
Melanophilharmostes	2606967	genus
- Melanophilharmostes carinatus	2903152	species
- Melanophilharmostes poggii	2607242	species
- Melanophilharmostes zicsii	2934990	species
unclassified Melanophilharmostes	2934991	no rank
Nesopalla	2606968	genus
- Nesopalla iviei	2607243	species
Perignamptus	2606971	genus
unclassified Perignamptus	2903166	no rank
Petrovitzostes	2606972	genus
- Petrovitzostes guineensis	2607244	species
Philharmostes	295526	genus
- Philharmostes basilewskyi	2607245	species
- Philharmostes grebennikovi	2607246	species
- Philharmostes interruptus	2607247	species
- Philharmostes ornatus	2607248	species
- Philharmostes pseudumbratilis	2607249	species
- Philharmostes werneri	2607250	species
unclassified Philharmostes	2607271	no rank
Pseudopterorthochaetes	2606973	genus
- Pseudopterorthochaetes demirei	2607251	species
- Pseudopterorthochaetes genierorum	2903153	species
- Pseudopterorthochaetes miomboincola	2903154	species
unclassified Pseudopterorthochaetes	2903168	no rank
Pseudosynarmostes	2606974	genus
- Pseudosynarmostes mitsinjo	2607252	species
Pterorthochaetes	1552080	genus
- Pterorthochaetes insularis	3085237	species
unclassified Pterorthochaetes	2622785	no rank
Synarmostes	2606977	genus
- Synarmostes humilis	2607253	species
- Synarmostes tibialis	2903151	species
unclassified Synarmostes	2903171	no rank
unclassified Ceratocanthinae	2606983	no rank
- Ceratocanthinae sp. BMNH 836004	1142399	species
Hybosorinae	2607020	subfamily
Brenskea	3085240	genus
- Brenskea coronata	3085241	species
Coilodes	3085242	genus
- Coilodes castaneus	3085243	species
Hybosorus	351681	genus
- Hybosorus illigeri	351689	species
- Hybosorus orientalis	2778438	species
- Hybosorus ruficornis	3085233	species
unclassified Hybosorus	2644960	no rank
Metachaetodus	3085244	genus
- Metachaetodus brunneicollis	3085245	species
Phaeochroops	1627862	genus
- Phaeochroops acuticollis	1627863	species
- Phaeochroops rattus	3085236	species
Phaeochrous	295538	genus
- Phaeochrous emarginatus	2891615	species
- Phaeochrous mashunus	295539	species
unclassified Phaeochrous	2621734	no rank
Liparochrinae	2607022	subfamily
Antiochrus	2903160	genus
- Antiochrus aberrans	2903161	species
Liparochrus	295536	genus
- Liparochrus infantus	2903158	species
- Liparochrus matthewsi	2903157	species
- Liparochrus sculptilis	295537	species
- Liparochrus silphoides	2903155	species
unclassified Liparochrus	2634099	no rank
Pachyplectrinae	2607021	subfamily
Pachyplectrus	1542265	genus
- Pachyplectrus laevis	1542266	species
unclassified Hybosoridae	1989351	no rank
- Hybosoridae sp.	2981733	species
- Hybosoridae sp. BIOUG04772-C01	2001343	species
- Hybosoridae sp. BIOUG04772-C03	1998240	species
- Hybosoridae sp. BIOUG04772-C11	1989354	species
- Hybosoridae sp. BIOUG04772-D01	1992314	species
- Hybosoridae sp. BIOUG04772-D07	1989352	species
- Hybosoridae sp. BIOUG04772-D10	1992313	species
- Hybosoridae sp. BIOUG04772-E11	2001342	species
- Hybosoridae sp. BIOUG05357-F01	1989355	species
- Hybosoridae sp. BIOUG05357-F02	1989353	species
- Hybosoridae sp. BIOUG18623-E12	1998239	species
- Hybosoridae sp. BOLD:AAP7884	2791707	species
- Hybosoridae sp. BOLD:AAP7958	2791708	species
- Hybosoridae sp. CNC COL EO119822	2917907	species
Lucanidae	41105	family	stag beetles
Aesalinae	614532	subfamily
Aesalini	614537	tribe
Aesalus	273956	genus
Himaloaesalus	2583517	genus
Nicagini	614539	tribe
Nicagus	273959	genus
Lampriminae	614533	subfamily
Lamprima	614624	genus
- Lamprima adolphinae	617909	species
- Lamprima aenea	617910	species
- Lamprima insularis	617913	species
- Lamprima latreillii	617914	species
unclassified Lamprima	2624452	no rank
Phalacrognathus	614625	genus
- Phalacrognathus muelleri	618351	species
Streptocerus	614626	genus
- Streptocerus speciosus	618668	species
Lucaninae	614534	subfamily
Aegognathus	614544	genus
- Aegognathus waterhousei	617265	species
Aegus	273935	genus
- Aegus chrysomelinus	617301	species
- Aegus currani	617309	species
- Aegus fukiensis	617332	species
- Aegus kurosawai	617372	species
Aegus laevicollis	273936	species
- Aegus nakaneorum	617399	species
- Aegus parallelus	617409	species
- Aegus philippinensis	618872	species
unclassified Aegus	2625008	no rank
Allotopus	614546	genus
- Allotopus moellenkampi	617479	species
- Allotopus rosenbergi	617480	species
Amneidus	614547	genus
- Amneidus godefroyi	617481	species
Apterocyclus	614550	genus
- Apterocyclus honoluluensis	617486	species
Apterodorcus	614551	genus
- Apterodorcus bacchus	617488	species
Auxicerus	614552	genus
- Auxicerus platyceps	617491	species
Cacostomus	614556	genus
- Cacostomus floralis	617498	species
- Cacostomus subvittata	617500	species
unclassified Cacostomus	2631510	no rank
Calcodes	614557	genus
- Calcodes aeratus	617501	species
- Calcodes svenjae	617518	species
Cantharolethrus	614558	genus
- Cantharolethrus steinheili	617527	species
Capreolucanus	614559	genus
- Capreolucanus sicardi	617528	species
unclassified Capreolucanus	3095665	no rank
Cardanus	614560	genus
- Cardanus sulcithorax	617537	species
Casignetus	614561	genus
- Casignetus spixi	617541	species
Chiasognathus	614563	genus
- Chiasognathus grantii	1627843	species
- Chiasognathus latreillei	617578	species
- Chiasognathus mniszechii	617579	species
- Chiasognathus sombrus	1627844	species
Chileistomus	1081463	genus
- Chileistomus cucullatus	1081464	species
Colophon	614565	genus
- Colophon cameroni	617584	species
Colophon eastmani	617586	species
- Colophon endroedyi	617587	species
- Colophon haughtoni	617588	species
- Colophon izardi	617589	species
- Colophon kawaii	617590	species
- Colophon montisatris	617591	species
- Colophon neli	617592	species
- Colophon primosi	617594	species
- Colophon stokoei	1380083	species
- Colophon westwoodi	617597	species
- Colophon whitei	617598	species
unclassified Colophon	2634973	no rank
Cyclommatus	614566	genus
- Cyclommatus alagari	617600	species
- Cyclommatus albersi	619068	species
Cyclommatus asahinai	617602	species
Cyclommatus assamensis	2906887	species
- Cyclommatus bicolor	617603	species
- Cyclommatus canaliculatus	617606	species
- Cyclommatus cupreonitens	617608	species
- Cyclommatus dehaani	617609	species
- Cyclommatus elaphus	617610	species
- Cyclommatus eximius	617612	species
- Cyclommatus imperator	617618	species
- Cyclommatus lunifer	617622	species
- Cyclommatus margaritae	617624	species
Cyclommatus metallifer	617626	species
- Cyclommatus mniszechi	617628	species
- Cyclommatus nagaii	2079320	species
- Cyclommatus pulchellus	617635	species
- Cyclommatus scutellaris	617638	species
Cyclommatus strigiceps	617644	species
- Cyclommatus suzumurai	617647	species
- Cyclommatus truncatus	2906891	species
- Cyclommatus weinreichi	617653	species
- Cyclommatus zuberi	617654	species
Dorculus	614571	genus
- Dorculus lombokensis	617662	species
Dorcus	41106	genus
- Dorcus cervulus	3074309	species
- Dorcus davidis	617670	species
- Dorcus gracilicornis	617671	species
- Dorcus hansi	2593008	species
- Dorcus hirticornis	3074310	species
- Dorcus koreanus	667726	species
- Dorcus linwenhsini	3074305	species
- Dorcus metacostatus	244526	species
Dorcus montivagus	231737	species
- Dorcus parallelipipedus	41107	species	small stag beetle; lesser stag beetle
- Dorcus parallelus	617680	species
- Dorcus ratiocinativus	617685	species
Dorcus rectus	231732	species
- Dorcus rubrofemoratus	231736	species
- Dorcus taiwanicus	348880	species
- Dorcus tanakai	3074306	species
- Dorcus tenuihirsutus	2651293	species
- Dorcus tityus	1627845	species
- Dorcus ursulus	2651294	species
Dynodorcus	614572	subgenus
Erichius	1081465	genus
- Erichius brevicollis	1081466	species
- Erichius caelatus	1081467	species
- Erichius calverti	1081468	species
- Erichius costatus	1081469	species
- Erichius darwinii	1081470	species
Erichius fairmairii	1081471	species
- Erichius fasciatus	1081474	species
- Erichius femoralis	1081475	species
- Erichius franzae	1081476	species
- Erichius virgatus	1081477	species
Erichius vittatus	1081478	species
- Eucarteria	1627859	genus
Eulepidius	614575	genus
- Eulepidius luridus	617712	species
Falcicornis	3074311	genus
- Falcicornis himalayae	3423382	species
- Falcicornis moellenkampi	3423383	species
- Falcicornis ruficrus	3423384	species
- Falcicornis taibaishanensis	3074312	species
unclassified Falcicornis	3423904	no rank
Figulus	273948	genus
- Figulus binodulus	273949	species
- Figulus boninensis	273950	species
- Figulus punctatus	509906	species
unclassified Figulus	2641846	no rank
Geodorcus	153170	genus
- Geodorcus capito	153196	species
- Geodorcus helmsi	153197	species
- Geodorcus novaezealandiae	1081458	species
- Geodorcus philpotti	617853	species
- Geodorcus sororum	1124697	species
unclassified Geodorcus	2644270	no rank
Gnaphaloryx	614577	genus
- Gnaphaloryx opacus	617859	species
Hemisodorcus	614579	genus
- Hemisodorcus arrowi	617863	species
- Hemisodorcus derelictus	617864	species
- Hemisodorcus donckieri	617865	species
- Hemisodorcus macleayii	3074307	species
Hemisodorcus rubrofemoratus	617871	species
- Hemisodorcus sinensis	617872	species
Hexarthrius	614581	genus
- Hexarthrius mandibularis	617882	species
- Hexarthrius vitalisi	617888	species
Homoderus	614582	genus
- Homoderus gladiator	617896	species
- Homoderus mellyi	617897	species
Kirchnerius	2738764	genus
- Kirchnerius guangxii	2738765	species
Leptinopterus	614585	genus
- Leptinopterus femoratus	617925	species
Lissotes	614587	genus
- Lissotes cancroides	617252	species
- Lissotes cornutus	617958	species
- Lissotes menalcas	617253	species
- Lissotes rudis	617976	species
- Lissotes subcaeruleus	617977	species
unclassified Lissotes	2623216	no rank
Lucanus	41108	genus
- Lucanus barbarossa	617985	species
- Lucanus boileaui	2683037	species
- Lucanus capreolus	617990	species
Lucanus cervus	41109	species
- Lucanus chengyuani	2735355	species
- Lucanus datunensis	617994	species
- Lucanus dongi	3087084	species
- Lucanus dybowskii	3238474	species
- Lucanus elaphus	618004	species
- Lucanus ferriei	273938	species
- Lucanus formosanus	618008	species
- Lucanus fortunei	618010	species
- Lucanus fryi	618012	species
- Lucanus fujianensis	2937465	species
- Lucanus hayashii	618019	species
- Lucanus hermani	618020	species
- Lucanus ibericus	618022	species
- Lucanus imitator	618023	species
Lucanus kanoi	618025	species
- Lucanus klapperichi	618029	species
Lucanus kraatzi	618031	species
- Lucanus kurosawai	618032	species
- Lucanus laetus	618033	species
Lucanus laminifer	618034	species
- Lucanus liuweii	3418562	species
- Lucanus liuyei	2952596	species
- Lucanus luci	3087085	species
- Lucanus macrophyllus	619152	species
Lucanus maculifemoratus	231745	species
- Lucanus mazama	590157	species	cottonwood stag beetle
- Lucanus miwai	618043	species
- Lucanus miwai complex sp. Miaoli taxon	2995623	species
- Lucanus ngheanus	3087086	species
Lucanus ogakii	665918	species
- Lucanus parryi	618050	species
- Lucanus placidus	618052	species
- Lucanus planeti	618053	species
- Lucanus prometheus	618055	species
- Lucanus smithii	618062	species
- Lucanus swinhoei	618064	species
Lucanus szetschuanicus	618065	species
Lucanus tetraodon	618066	species
Lucanus thibetanus	618068	species
- Lucanus treadon	1142411	species
- Lucanus wuyishanensis	618075	species
- Lucanus yulaoensis	2819904	species
unclassified Lucanus	2646290	no rank
Macrodorcas	614589	genus
- Macrodorcas cuongi	3087020	species
- Macrodorcas mochizukii	618098	species
- Macrodorcas negrei	618101	species
- Macrodorcas seguyi	618109	species
Macrodorcas striatipennis	231734	species
Mesotopus	614590	genus
- Mesotopus regius	1627847	species
Mesotopus tarandus	618116	species
Metallactulus	614593	genus
- Metallactulus parvulus	618140	species
unclassified Metallactulus	2843189	no rank
Neolucanus	273939	genus
- Neolucanus armatus	618151	species
- Neolucanus baladeva	618152	species
- Neolucanus castanopterus	618156	species
- Neolucanus cf. doro NCHU:YP07	1432624	species
- Neolucanus cf. doro NCHU:YP08	1432625	species
- Neolucanus cf. doro NCHU:YP09	1432626	species
- Neolucanus cf. doro YP01	1262367	species
- Neolucanus cf. doro YP02	1262368	species
- Neolucanus cf. doro YP03	1262369	species
- Neolucanus cf. doro YP04	1262370	species
- Neolucanus cf. doro YP05	1262371	species
- Neolucanus cf. doro YP06	1262372	species
- Neolucanus cf. doro YP07	1262373	species
- Neolucanus cf. doro YP08	1262374	species
- Neolucanus cf. doro YP09	1262375	species
- Neolucanus donckieri	618163	species
Neolucanus doro	618164	species
- Neolucanus eugeniae	618165	species
- Neolucanus hagiangensis	3087098	species
Neolucanus maximus	618182	species
Neolucanus nitidus	618184	species
- Neolucanus oberthuri	618185	species
- Neolucanus parryi	618189	species
- Neolucanus perarmatus	618191	species
Neolucanus protogenetivus	273940	species
- Neolucanus pseudopacus	3087099	species
- Neolucanus punctulatus	3087100	species
- Neolucanus robustus	3383346	species
- Neolucanus rufus	618192	species
- Neolucanus rutilans	618193	species
Neolucanus sinicus	618197	species
- Neolucanus swinhoei	618200	species
- Neolucanus tao	618202	species
- Neolucanus vicinus	618203	species
unclassified Neolucanus	3071901	no rank
Nigidionus	614595	genus
- Nigidionus parryi	618206	species
Nigidius	273951	genus
- Nigidius acutangulus	618207	species
- Nigidius baeri	618210	species
- Nigidius delegorguei	618223	species
- Nigidius formosanus	618230	species
- Nigidius lewisi	273952	species
- Nigidius miwai	618256	species
- Nigidius sinicus	2950530	species
unclassified Nigidius	2646643	no rank
Noseolucanus	618279	genus
- Noseolucanus denticulus	1627848	species
- Noseolucanus rugosus	618280	species
Odontolabis	614597	genus
Odontolabis cuvera	618294	species
Odontolabis mouhoti	618316	species
Odontolabis platynota	618320	species
Odontolabis siva	618322	species
- Odontolabis somneri	618324	species
Platyceroides	614529	genus
unclassified Platyceroides	2648038	no rank
Platyceropsis	614531	genus
- Platyceropsis keeni	614635	species
Platycerus	273943	genus
Platycerus acuticollis	509907	species
- Platycerus akitaorum	509903	species
Platycerus albisomni	623701	species
- Platycerus auriceps	2678544	species
- Platycerus bashanicus	618359	species
- Platycerus businskyi	614511	species
- Platycerus canae	2678545	species
- Platycerus caprea	614637	species
- Platycerus caraboides	614512	species
- Platycerus caucasicus	614513	species
Platycerus consimilis	614514	species
- Platycerus cupreimicans	614515	species
- Platycerus cyanidraconis	623700	species
Platycerus delicatulus	273944	species
- Platycerus depressus	1895299	species
- Platycerus diluvialis	2678546	species
- Platycerus dundai	614516	species
- Platycerus feminatus	614517	species
- Platycerus hiurai	614518	species
Platycerus hongwonpyoi	614519	species
- Platycerus kawadai	509899	species
- Platycerus kitawakii	618363	species
- Platycerus ladyae	614521	species
- Platycerus mandibularis	2678548	species
- Platycerus masumotoi	2678549	species
- Platycerus nagahatai	623697	species
- Platycerus oregonensis	614522	species
- Platycerus primigenius	618368	species
- Platycerus quercus	3393976	species
Platycerus rugosus	614523	species
- Platycerus spinifer	618372	species
- Platycerus sue	618373	species
- Platycerus sugitai	509904	species
Platycerus tabanai	614524	species
Platycerus takakuwai	623702	species
- Platycerus tangi	623699	species
- Platycerus tieguanzi	614525	species
Platycerus turnai	614526	species
- Platycerus urushiyamai	509905	species
- Platycerus virescens	614527	species
Platycerus viridicuprus	623703	species
- Platycerus xiongmao	2678552	species
- Platycerus yangi	2678553	species
- Platycerus yeren	623698	species
- Platycerus yingqii	2678554	species
unclassified Platycerus	2649287	no rank
Prismognathus	231742	genus
Prismognathus angularis	231743	species
- Prismognathus dauricus	273934	species
Prismognathus davidis	618379	species
- Prismognathus formosanus	618381	species
- Prismognathus piluensis	618393	species
- Prismognathus prossi	618395	species
- Prismognathus triapicalis	2906892	species
Prosopocoilus	231739	genus
Prosopocoilus astacoides	618406	species
Prosopocoilus bulbosus	618416	species
- Prosopocoilus confucius	618424	species
Prosopocoilus forficula	618451	species
- Prosopocoilus formosanus	1706339	species
- Prosopocoilus girafa	618460	species
- Prosopocoilus gracilis	1672220	species
Prosopocoilus inclinatus	231740	species
- Prosopocoilus inquinatus	618466	species
Prosopocoilus laterotarsus	618480	species
- Prosopocoilus motschulskyi	618490	species
- Prosopocoilus natalensis	618493	species
- Prosopocoilus pseudodissimilis	273933	species
- Prosopocoilus reni	2588723	species
- Prosopocoilus savagei	1627851	species
- Prosopocoilus sericeus	618509	species
- Prosopocoilus spineus	618515	species
- Prosopocoilus suturalis	618518	species
unclassified Prosopocoilus	2647816	no rank
Pseudorhaetus	614606	genus
- Pseudorhaetus oberthuri	618540	species
Pseudorhaetus sinicus	618541	species
Pycnosiphorus	614607	genus
Pycnosiphorus lessonii	618559	species
- Pycnosiphorus philippi	618564	species
Rhaetulus	273931	genus
Rhaetulus crenatus	618569	species
- Rhaetulus didieri	618570	species
- Rhaetulus recticornis	273932	species
Rhaetus	614608	genus
- Rhaetus westwoodii	618572	species
Ryssonotus	614609	genus
- Ryssonotus laticeps	618578	species
- Ryssonotus nebulosus	1627852	species
Serrognathus	614612	genus
- Serrognathus alcides	618607	species
- Serrognathus bucephalus	618610	species
- Serrognathus castanicolor	3074308	species
- Serrognathus consentaneus	2891608	species
- Serrognathus eurycephalus	618615	species
- Serrognathus kyanrauensis	618616	species
- Serrognathus miwai	618621	species
- Serrognathus platymelus	618622	species
Serrognathus reichei	618624	species
Serrognathus titanus	618631	species
unclassified Serrognathus	2647554	no rank
Sphaenognathus	614613	genus
- Sphaenognathus alticollis	618639	species
- Sphaenognathus giganteus	618648	species
- Sphaenognathus kolbei	618650	species
- Sphaenognathus metallifer	618654	species
- Sphaenognathus monguilloni	618655	species
- Sphaenognathus munchowae	618656	species
- Sphaenognathus nobilis	618657	species
- Sphaenognathus oberon	618658	species
- Sphaenognathus queenslandicus	618662	species
- Sphaenognathus taschenbergi	618666	species
- Sphaenognathus xerophilus	618667	species
Velutinodorcus	614617	genus
- Velutinodorcus carinulatus	618692	species
- Velutinodorcus japonicus	618693	species
- Velutinodorcus velutinus	618696	species
Weinreichius	614619	genus
- Weinreichius perroti	618698	species
Xiphodontus	614620	genus
- Xiphodontus antilope	618699	species
Yumikoi	614621	genus
- Yumikoi makii	618702	species
Syndesinae	614536	subfamily
Ceruchus	219418	genus
- Ceruchus chrysomelinus	617560	species
Ceruchus lignarius	273953	species
- Ceruchus minor	617564	species
- Ceruchus piceus	617566	species
- Ceruchus striatus	617570	species
unclassified Ceruchus	2638017	no rank
Sinodendron	614542	genus
- Sinodendron cylindricum	618634	species
- Sinodendron rugosum	618636	species
- Sinodendron yunnanense	618637	species
- unclassified Lucanidae	614530	no rank
Ochodaeidae	110055	family
Cucochodaeus	702047	genus
- Cucochodaeus sparsus	702048	species
Ochodaeus	295541	genus
- Ochodaeus cornifrons	1542175	species
- Ochodaeus montanus	1142412	species
- Ochodaeus solskyi	1627849	species
unclassified Ochodaeus	2624517	no rank
- Ochodaeus sp. BM678423	397610	species
- Ochodaeus sp. BMNH 678422	465214	species
- Ochodaeus sp. TJH-2004	295542	species
Parochodaeus	2680953	genus
- Parochodaeus biarmatus	2680954	species
Xenochodaeus	1710482	genus
- Xenochodaeus musculus	2109466	species
- Xenochodaeus simplex	1710483	species
Passalidae	110046	family
Aceraius	459810	genus
- Aceraius grandis	1453159	species
- Aceraius helferi	459811	species
unclassified Aceraius	2642673	no rank
- Aceraius sp. INB096	2055317	species
- Aceraius sp. SIK-2015	1627853	species
Ameripassalus	2652949	genus
- Ameripassalus guatemalensis	2652950	species
Analaches	1269065	genus
- Analaches australiensis	1269066	species
Arrox	2652951	genus
- Arrox agassizi	2652952	species
Aulacocyclus	295543	genus
- Aulacocyclus edentulus	459791	species
- Aulacocyclus fracticornis	1269060	species
- Aulacocyclus kaupi	1542143	species
- Aulacocyclus rouxi	459789	species
- Aulacocyclus tambourinensis	2652914	species
- Aulacocyclus tricuspis	459790	species
unclassified Aulacocyclus	2648415	no rank
- Aulacocyclus sp. 1 HU-2012	1269062	species
- Aulacocyclus sp. 2 HU-2012	1269063	species
- Aulacocyclus sp. 3 HU-2012	1269064	species
- Aulacocyclus sp. BMNH 670848	465182	species
- Aulacocyclus sp. DDM_3296	2652985	species
- Aulacocyclus sp. NLG-2014	1551871	species
- Aulacocyclus sp. TJH-2004	295544	species
Ceracupes	459804	genus
- Ceracupes arrowi	459807	species
- Ceracupes fronticornis	459805	species
- Ceracupes yui	459806	species
unclassified Ceracupes	2639352	no rank
- Ceracupes sp. INB057	2055352	species
Chondrocephalus	1112829	genus
- Chondrocephalus gemmae	2652915	species
- Chondrocephalus granulifrons	2583727	species
- Chondrocephalus purulensis	1112830	species
- Chondrocephalus salvadorae	2025040	species
unclassified Chondrocephalus	2652986	no rank
- Chondrocephalus sp. 1 CFBB-2020	2755229	species
- Chondrocephalus sp. 2 CFBB-2020	2755230	species
- Chondrocephalus sp. DDM_3228	2652987	species
- Chondrocephalus sp. DDM_3241	2652988	species
Comacupes	459798	genus
- Comacupes cavicornis	459802	species
- Comacupes cylindraceus	459801	species
- Comacupes masoni	459803	species
- Comacupes stoliczkae	459799	species
Cylindrocaulus	273923	genus
- Cylindrocaulus bucerus	459787	species
- Cylindrocaulus davidi	459788	species
- Cylindrocaulus patalis	273924	species
Erionomus	2755233	genus
- Erionomus planiceps	2755234	species
Gonatas	1552018	genus
unclassified Gonatas	2634018	no rank
- Gonatas sp. NLG-2014	1552019	species
Heliscus	2025064	genus
- Heliscus eclipticus	2652916	species
- Heliscus tropicus	2025065	species
- Heliscus vazquezae	2025066	species
Leptaulax	459812	genus
- Leptaulax bicolor	459813	species
- Leptaulax dentatus	1542162	species
- Leptaulax formosanus	1689726	species
- Leptaulax koreanus	2607329	species
Mastachilus	1269067	genus
- Mastachilus quaestionis	1269069	species
unclassified Mastachilus	2628196	no rank
- Mastachilus sp. HU-2012	1269070	species
- Mastachilus sp. NLG-2014	1551921	species
- Mastachilus sp. n. 1 HU-2012	1269068	species
Odontotaenius	295545	genus
- Odontotaenius cryptodisjunctus	3118357	species
- Odontotaenius disjunctus	295546	species	bess beetle; betsy beetle; patent leather beetle
- Odontotaenius floridanus	3118356	species
- Odontotaenius jackmanicus	3118358	species
- Odontotaenius striatopunctatus	2025041	species
- Odontotaenius zodiacus	2025042	species
unclassified Odontotaenius	2652990	no rank
- Odontotaenius sp. 1 CFBB-2020	2755470	species
- Odontotaenius sp. 2 CFBB-2020	2755471	species
- Odontotaenius sp. DDM_3190	2652991	species
- Odontotaenius sp. DDM_3196	2652992	species
Ogyges	1115857	genus
- Ogyges aluxi	2652917	species
- Ogyges championi	1115858	species
- Ogyges crassulus	2652918	species
- Ogyges laevissimus	1148777	species
- Ogyges tzutuhili	2583728	species
Oileus	1115859	genus
- Oileus bifidus	2025043	species
- Oileus gasparilomi	2652919	species
- Oileus rimator	2583729	species
- Oileus sargi	1115860	species
unclassified Oileus	2652993	no rank
- Oileus sp. DDM_3247	2652994	species
Ophrygonius	2897801	genus
unclassified Ophrygonius	2897802	no rank
- Ophrygonius sp.	2897803	species
Passalus	459808	genus
- Passalus abortivus	2652920	species
- Passalus alfaroi	2652921	species
- Passalus bucki	2652922	species
- Passalus coniferus	2652925	species
- Passalus convexus	2583731	species
- Passalus cuneatus	2728599	species
- Passalus curtus	2652926	species
- Passalus gaboi	3055481	species
- Passalus inops	2652927	species
- Passalus interruptus	2583730	species
- Passalus interstitialis	1115856	species
- Passalus jansoni	2652928	species
- Passalus lunaris	2652929	species
- Passalus paucuvillosus	3055253	species
- Passalus punctatostriatus	2211546	species
- Passalus punctiger	1112828	species
- Passalus ruehli	2652930	species
- Passalus rugosus	2652932	species
- Passalus serankuai	3055254	species
- Passalus unimagdalenae	3055255	species
unclassified Passalus	2620060	no rank
- Passalus sp. 14YB	2728657	species
- Passalus sp. 15YB	2728658	species
- Passalus sp. 16YB	2728659	species
- Passalus sp. 3YB	2728660	species
- Passalus sp. Pa030625	459809	species
- Passalus sp. SIK-2015	1627855	species
Passipassalus	2652953	genus
- Passipassalus nukak	2652954	species
Paxillus	2583738	genus
- Paxillus leachii	2583739	species
Petrejoides	1564290	genus
- Petrejoides chiapasae	2025045	species
- Petrejoides guatemalae	2025046	species
- Petrejoides haagi	2652934	species
- Petrejoides jalapensis	2025047	species
- Petrejoides orizabae	2025048	species
- Petrejoides tenuis	2025049	species
unclassified Petrejoides	2652995	no rank
- Petrejoides sp. 1 CFBB-2020	2755224	species
- Petrejoides sp. 1YB	2728723	species
- Petrejoides sp. 2 CFBB-2020	2755225	species
- Petrejoides sp. 3 CFBB-2020	2755472	species
- Petrejoides sp. DDM_3208	2652996	species
- Petrejoides sp. DDM_3246	2652997	species
- Petrejoides sp. DDM_3248	2652998	species
Pharochilus	2652955	genus
- Pharochilus nitidulus	2652956	species
- Pharochilus politus	2652957	species
Popilius	1148778	genus
- Popilius erotylus	2025050	species
- Popilius gibbosus	2652935	species
- Popilius marginatus	2652936	species
unclassified Popilius	2728663	no rank
- Popilius sp. 1YB	2728664	species
Proculejus	2170619	genus
unclassified Proculejus	2652999	no rank
- Proculejus sp. 1 CFBB-2020	2755226	species
- Proculejus sp. 2 CFBB-2020	2755227	species
- Proculejus sp. 3 CFBB-2020	2755228	species
- Proculejus sp. DDM_2867	2653000	species
- Proculejus sp. DDM_3195	2653001	species
- Proculejus sp. DDM_3199	2653002	species
- Proculejus sp. DDM_3211	2653003	species
Proculus	459814	genus
- Proculus goryi	2652937	species
- Proculus mniszechi	459815	species
Pseudacanthus	2583736	genus
- Pseudacanthus aztecus	2583737	species
- Pseudacanthus junctistriatus	2652938	species
- Pseudacanthus subopacus	2652939	species
Pseudoarrox	2652958	genus
- Pseudoarrox caldasi	2652959	species
Ptichopus	2652960	genus
- Ptichopus angulatus	2652961	species
Publius	1882557	genus
unclassified Publius	2728665	no rank
- Publius sp. 1YB	2728666	species
Rhodocanthopus	3055258	genus
- Rhodocanthopus caelatus	3070647	species
- Rhodocanthopus clypeoneleus	3070649	species
- Rhodocanthopus cognatus	3070650	species
- Rhodocanthopus maillei	3055259	species
- Rhodocanthopus rufiventris	3055260	species
- Rhodocanthopus spiniger	3070651	species
Spurius	2025052	genus
- Spurius bicornis	2025053	species
unclassified Spurius	2627524	no rank
- Spurius sp. CB049	2025062	species
- Spurius sp. CB050	2025063	species
Taeniocerus	459792	genus
- Taeniocerus bicanthatus	459796	species
- Taeniocerus mourzinei	459797	species
- Taeniocerus platypus	459793	species
- Taeniocerus pygmaeus	459794	species
unclassified Taeniocerus	2632562	no rank
- Taeniocerus sp. TU980920	459795	species
Tonantzin	2709219	genus
- Tonantzin tepetl	2583740	species
Undulifer	2652962	genus
- Undulifer incisus	2652963	species
- Undulifer nigidioides	2652964	species
Verres	1115861	genus
- Verres cavicollis	2583732	species
- Verres corticicola	1564294	species
- Verres deficiens	2652940	species
- Verres furcilabris	2583733	species
- Verres hageni	1115862	species
- Verres onorei	2652941	species
- Verres sternbergianus	2728625	species
unclassified Verres	2640103	no rank
- Verres sp. SIK-2015	1627856	species
Veturius	1564292	genus
- Veturius aspina	2652942	species
- Veturius assimilis	2583734	species
- Veturius cirratus	2755222	species
- Veturius crassus	2755223	species
- Veturius impressus	3055256	species
- Veturius oberthuri	2583735	species
- Veturius standfussi	3055257	species
- Veturius talamacaensis	2652943	species
- Veturius tuberculifrons	2652944	species
unclassified Veturius	2653004	no rank
- Veturius sp. 1YB	2728724	species
- Veturius sp. DDM_3201	2653005	species
Vindex	1112832	genus
- Vindex agnoscendus	2652945	species
- Vindex gonzaloi	2652946	species
- Vindex sculptilis	2652947	species
unclassified Vindex	2637328	no rank
- Vindex sp. 1 CFBB-2020	2755231	species
- Vindex sp. 2 CFBB-2020	2755232	species
- Vindex sp. CB015	2025051	species
- Vindex sp. DDM_3250	2653006	species
Xylopassaloides	1148780	genus
- Xylopassaloides chortii	1148781	species
Yumtaax	2025054	genus
- Yumtaax cameliae	2025059	species
- Yumtaax imbellis	2025055	species
- Yumtaax jimenezi	2025060	species
- Yumtaax laticornis	2025056	species
- Yumtaax mazatecus	2025057	species
- Yumtaax recticornis	2025058	species
- Yumtaax veracrucensis	2025061	species
unclassified Passalidae	1205570	no rank
- Passalidae sp. BOLD:AAG2092	2660200	species
- Passalidae sp. GENSP01	1205571	species
- Passalidae sp. GENSP02	1205572	species
- Proculini sp. DDM_3251	2653007	species
Pleocomidae	110050	family
Acoma	1542200	genus
- Acoma glabrata	1542201	species
Pleocoma	295547	genus
- Pleocoma australis	1542188	species
- Pleocoma rubiginosa	558336	species
unclassified Pleocoma	2627975	no rank
- Pleocoma sp. TH-2004-5	295548	species
Scarabaeidae	7055	family	lamellicorn beetles; scarab beetles
Aclopinae	702309	subfamily
Aclopus	702310	genus
unclassified Aclopus	2628989	no rank
Desertaclopus	2570516	genus
- Desertaclopus lucasi	2570517	species
Gracilaclopus	2570518	genus
- Gracilaclopus electricus	2570519	species
Neophaenognatha	702814	genus
- Neophaenognatha jenseni	702815	species
Phaenognatha	1552068	genus
unclassified Phaenognatha	2648062	no rank
Allidiostomatinae	702055	subfamily
Allidiostoma	702045	genus
- Allidiostoma hirtum	702046	species
- Allidiostoma landbecki	2570512	species
- Allidiostoma strobeli	2570513	species
unclassified Allidiostoma	2639033	no rank
Aphodiinae	166306	subfamily
Acrossidius	207111	genus
- Acrossidius tasmaniae	207112	species
unclassified Acrossidius	2624899	no rank
Acrossus	225234	genus
- Acrossus atratus	3228759	species
- Acrossus carpetanus	207133	species
- Acrossus depressus	207143	species
- Acrossus humerospinosus	3228760	species
- Acrossus luridus	207177	species
- Acrossus rufipes	207206	species
- Acrossus superatratus	3078809	species
unclassified Acrossus	3078807	no rank
Aegialia	206942	genus
- Aegialia arenaria	206943	species
- Aegialia crassa	207238	species
- Aegialia criddlei	1943727	species
- Aegialia cylindrica	2069338	species
- Aegialia lacustris	1483802	species
- Aegialia mimica	1971767	species
- Aegialia sabuleti	1553035	species
- Aegialia terminalis	1709555	species
unclassified Aegialia	2619709	no rank
Afrodiastictus	207260	genus
unclassified Afrodiastictus	2622582	no rank
Airapus	207367	genus
- Airapus henriettae	207368	species
unclassified Airapus	2638406	no rank
Alloblackburneus	2070432	genus
- Alloblackburneus lentus	2546194	species
Amerisaprus	3397864	genus
- Amerisaprus valdivia	3397868	species
unclassified Amerisaprus	3397881	no rank
Aphodius	166307	genus
Acanthobodilus	225232	subgenus
Adeloparius	465091	subgenus
Aganocrossus	225235	subgenus
Agoliinus	225245	subgenus
Agolius	225246	subgenus
Agrilinus	225247	subgenus
Alocoderus	225248	subgenus
Amidorus	225250	subgenus
Ammoecius	225251	subgenus
Anomius	225252	subgenus
Aphodaulacus	225572	subgenus
Aphodius	225253	subgenus
Aphodius incertae sedis	1552118	no rank
Biralus	225254	subgenus
Blackburneus	225547	subgenus
Bodilopsis	3016547	subgenus
Bodilus	225255	subgenus
Calamosternus	225256	subgenus
Chilothorax	225257	subgenus
Cinacanthus	225258	subgenus
Colobopterus	225259	subgenus
Coprimorphus	225260	subgenus
Esymus	225262	subgenus
Eudolus	225263	subgenus
Euorodalus	225264	subgenus
Eupleurus	225265	subgenus
Labarrus	225266	subgenus
Limarus	225267	subgenus
Liothorax	225268	subgenus
Loboparius	225550	subgenus
Loraphodius	225269	subgenus
Melaphodius	225273	subgenus
Mendidaphodius	225275	subgenus
Mesontoplatys	225549	subgenus
Nialaphodius	465099	subgenus
Nialus	225277	subgenus
Oromus	225281	subgenus
Otophorus	225282	subgenus
Paradidactylia	225576	subgenus
Parammoecius	225283	subgenus
Paremadus	225554	subgenus
Phaeaphodius	225284	subgenus
Phalacronothus	225285	subgenus
Plagiogonus	225287	subgenus
Planolinus	225288	subgenus
Platyderides	225546	subgenus
Pleuraphodius	225555	subgenus
Sigorus	225291	subgenus
Sinodiapterna	225575	subgenus
Stenotothorax	225573	subgenus
Subrinus	225292	subgenus
Sugrames	377214	subgenus
Teuchestes	225293	subgenus
Trichaphodius	225548	subgenus
Volinus	225319	subgenus
unclassified Aphodius	225295	no rank
Aphodopsammobius	1551970	genus
unclassified Aphodopsammobius	2641985	no rank
Ataenius	207344	genus
- Ataenius imbricatus	489792	species
- Ataenius monteithi	207373	species
- Ataenius opatroides	1925786	species
- Ataenius picinus	3375601	species
- Ataenius platensis	2680150	species
- Ataenius sciurus	207374	species
- Ataenius spretulus	1709596	species
- Ataenius strigatus	207375	species
unclassified Ataenius	2638424	no rank
Australammoecius	207376	genus
- Australammoecius occidentalis	207377	species
unclassified Australammoecius	2639125	no rank
Australaphodius	207262	genus
unclassified Australaphodius	2685188	no rank
Australoxenella	207378	genus
- Australoxenella conccina	207379	species
Brindalus	207380	genus
- Brindalus porcicollis	207381	species
Caelius	1944056	genus
- Caelius browni	1944057	species
- Caelius montanus	1944058	species
Caligodorus	1709651	genus
- Caligodorus opacus	1709652	species
Chiron	207382	genus
- Chiron cylindricus	207383	species
unclassified Chiron	2643996	no rank
Coptochirus	207346	genus
- Coptochirus n. sp. COPT_G	207347	species
- Coptochirus perlucidus	465120	species
Cryptoscatomaseter	2083760	genus
- Cryptoscatomaseter punctissimus	2083761	species
Dellacasiellus	2680579	genus
- Dellacasiellus kirni	2680580	species
Dialytellus	377169	genus
- Dialytellus tragicus	377190	species
unclassified Dialytellus	3054161	no rank
Dialytes	207227	genus
- Dialytes striatulus	207228	species
- Dialytes truncatus	207229	species
unclassified Dialytes	2642848	no rank
Diapterna	1710456	genus
- Diapterna omissa	1710457	species
Didactylia	207384	genus
- Didactylia parcus	207385	species
Drepanocanthoides	465224	genus
- Drepanocanthoides walshi	465225	species
Erezamus	465232	genus
- Erezamus unistriatus	1142402	species
unclassified Erezamus	2685573	no rank
Geomyphilus	1636549	genus
- Geomyphilus barrerai	1636551	species
- Geomyphilus pierai	1636550	species
unclassified Geomyphilus	2633762	no rank
Goiginus	465243	genus
unclassified Goiginus	2644798	no rank
Harmogaster	207386	genus
- Harmogaster exarata	207387	species
unclassified Harmogaster	2624533	no rank
Heptaulacus	225320	genus
- Heptaulacus testudinarius	225321	species
- Leiopsammodius	207388	genus
Lorditomaeus	207230	genus
- Lorditomaeus invenustus	207231	species
Martineziella	377171	genus
- Martineziella dutertrei	377198	species
Melinopterus	225274	genus
- Melinopterus consputus	207139	species
- Melinopterus prodromus	1971315	species
- Melinopterus punctatosulcatus	1501853	species
- Melinopterus sphacelatus	207110	species
- Melinopterus tingens	207218	species
unclassified Melinopterus	2908784	no rank
Neocalaphodius	3303172	genus
- Neocalaphodius moestus	3303174	species
Neoemadiellus	767848	genus
- Neoemadiellus humerosanguineum	767849	species
unclassified Neoemadiellus	2618612	no rank
Neoheptaulacus	1142441	genus
unclassified Neoheptaulacus	2631427	no rank
Neotrichiorhyssemus	1902231	genus
- Neotrichiorhyssemus hirsutus	1902232	species
Neotrichonotulus	1636552	genus
- Neotrichonotulus perotensis	1636553	species
Nimbus	225278	genus
- Nimbus affinis	3402512	species
- Nimbus contaminatus	3402504	species
- Nimbus johnsoni	3402513	species
- Nimbus obliteratus	3402514	species
- Nimbus proximus	225318	species
Nobiellus	2869604	genus
- Nobiellus bonnairei	207127	species
Nobius	225279	genus
- Nobius hepaticus	2867989	species
unclassified Nobius	3382381	no rank
Oscarinus	1710675	genus
- Oscarinus rusicola	1710676	species
Oxyomus	207232	genus
- Oxyomus sylvestris	207233	species
Pharaphodius	225286	genus
- Pharaphodius fiechteri	3382449	species
- Pharaphodius impurus	2868059	species
- Pharaphodius infinitus	3382457	species
- Pharaphodius marginellus	207180	species
- Pharaphodius rubricosus	207205	species
Planolinoides	1944085	genus
- Planolinoides borealis	1944086	species
Platytomus	207391	genus
- Platytomus caelicollis	207392	species
Pleurophorus	207393	genus
- Pleurophorus caesus	207394	species
unclassified Pleurophorus	2641994	no rank
Podotenus	207234	genus
- Podotenus fulviventris	1925791	species
- Podotenus storeyi	207235	species
unclassified Podotenus	2621525	no rank
Psammodius	464793	genus
- Psammodius asper	1553686	species
- Psammodius convexus	1569674	species
- Psammodius laevipennis	2340033	species
- Psammodius pierottii	2567391	species
Pseudoxyomus	207287	genus
unclassified Pseudoxyomus	2641893	no rank
Rhyparus	207395	genus
- Rhyparus australiae	207396	species
- Rhyparus opacus	1542192	species
unclassified Rhyparus	2628062	no rank
Rhyssemus	225574	genus
- Rhyssemus germanus	1588430	species
- Rhyssemus inscitus	3127446	species
- Rhyssemus ritsemae	207204	species
unclassified Rhyssemus	2834716	no rank
Saprosites	207397	genus
- Saprosites catenatus	207398	species
- Saprosites mendax	465152	species
unclassified Saprosites	2618554	no rank
Sybax	207236	genus
- Sybax impressicollis	1142423	species
- Sybax sulcicollis	207237	species
Tesarius	207399	genus
- Tesarius caelatus	207400	species
Trichiorhyssemus	1885631	genus
- Trichiorhyssemus asperulus	1885632	species
Trichonotulus	225294	genus
- Trichonotulus scrofa	207209	species
unclassified Aphodiinae	1105441	no rank
- Aphodiinae sp.	3067624	species
- Aphodiinae sp. AFX9206	3382355	species
- Aphodiinae sp. AFX9226	3382356	species
- Aphodiinae sp. AGA5675	3382357	species
- Aphodiinae sp. AGA5676	3382358	species
- Aphodiinae sp. AGA5678	3382359	species
- Aphodiinae sp. AGA5688	3382360	species
- Aphodiinae sp. AGA5693	3382361	species
- Aphodiinae sp. AGA5697	3382362	species
- Aphodiinae sp. AGA5698	3382363	species
- Aphodiinae sp. BOLD:AAH0197	2657175	species
- Aphodiinae sp. BOLD:AAH0198	2657172	species
- Aphodiinae sp. BOLD:AAH0464	2657173	species
- Aphodiinae sp. BOLD:AAN5996	1105442	species
- Aphodiinae sp. BOLD:AAP9995	2657174	species
Cetoniinae	7056	subfamily	flower beetles; goliath beetles; goliath beetles & flower beetles
Aethiessa	1207111	genus
- Aethiessa floralis	1207112	species
unclassified Aethiessa	2676944	no rank
Agestrata	1208515	genus
unclassified Agestrata	2634384	no rank
Anelaphinis	1207115	genus
- Anelaphinis simillima	1207116	species
Anoxia	1207117	genus
- Anoxia cypria	2563622	species
- Anoxia orientalis	3230646	species
- Anoxia pilosa	1476578	species
- Anoxia sardoa	1207118	species
- Anoxia villosa	1476579	species
unclassified Anoxia	2618981	no rank
Anthracophora	1163333	genus
- Anthracophora crucifera	3402269	species
- Anthracophora rusticola	1163330	species
Aphanesthes	2740238	genus
- Aphanesthes pullata	2740239	species
- Aphanesthes succinea	2740242	species
unclassified Aphanesthes	2740240	no rank
Apocnosis	1207165	genus
- Apocnosis striata	1207166	species
Argyropheges	1207113	genus
- Argyropheges kolbei	1207114	species
Atrichelaphinis	1207119	genus
- Atrichelaphinis nigropunctulata	1207120	species
- Atrichelaphinis tigrina	1207167	species
Atrichiana	1207168	genus
- Atrichiana placida	1207169	species
Bisallardiana	1551978	genus
- Bisallardiana decorticata	2740187	species
- Bisallardiana gymnopleura	2740188	species
- Bisallardiana variabilis	2740196	species
unclassified Bisallardiana	2637791	no rank
Campsiura	1207170	genus
- Campsiura javanica	1207171	species
- Campsiura mirabilis	3127969	species
Cetonia	290678	genus
Cetonia aurata	290679	species
- Cetonia aurataeformis	1207078	species
- Cetonia carthami	702042	species
- Cetonia delagrangei	1207085	species
- Cetonia magnifica	1618296	species
- Cetonia pilifera	2898101	species
- Cetonia viridiopaca	1618297	species
unclassified Cetonia	2627745	no rank
Chiloloba	383510	genus
- Chiloloba acuta	383520	species
- Chiloloba orientalis	2749413	species
Chlorobapta	1551988	genus
- Chlorobapta frontalis	2740197	species
- Chlorobapta goerlingi	2740198	species
unclassified Chlorobapta	2621221	no rank
Chondropyga	1551990	genus
- Chondropyga dorsalis	2740200	species
- Chondropyga insignicosta	2740201	species
unclassified Chondropyga	2639533	no rank
Chtonobius	1208518	genus
unclassified Chtonobius	2631683	no rank
Clinteria	2780918	genus
- Clinteria klugi	2780919	species
Clinterocera	2859702	genus
- Clinterocera nigra	2921222	species
- Clinterocera obsoleta	2859703	species
Clinteroides	1207174	genus
- Clinteroides permutans	1207175	species
Clithria	2740243	genus
- Clithria albersi	2740244	species
- Clithria eucnemis	2740245	species
- Clithria tibiale	2740246	species
Coenochilus	1208520	genus
- Coenochilus striatus	2962661	species
unclassified Coenochilus	2621974	no rank
Coilodera	1207127	genus
- Coilodera penicillata	1207128	species
Comythovalgus	1208522	genus
unclassified Comythovalgus	2649628	no rank
Conradtia	1207176	genus
- Conradtia principalis	1207177	species
Coptomia	1207129	genus
- Coptomia laevis	1207130	species
Cosmiomorpha	3087145	genus
- Cosmiomorpha pacholatkoi	3087146	species
Cotinis	1248212	genus
- Cotinis mutabilis	1248213	species
- Cotinis nitida	1804801	species	green June beetle
Cremastocheilus	302189	genus
unclassified Cremastocheilus	2641318	no rank
Cymophorus	1207131	genus
- Cymophorus cf. leucostictus DA-2012	1207323	species
unclassified Cymophorus	2677122	no rank
Cyriodera	1207133	genus
- Cyriodera tuberculicollis	1207134	species
Cyrtothyrea	1207135	genus
- Cyrtothyrea cf. marginalis DA-2012	1207325	species
- Cyrtothyrea testaceoguttata	1207137	species
Dasyvalgus	2055606	genus
unclassified Dasyvalgus	2628780	no rank
Diaphonia	2740247	genus
- Diaphonia bacchusi	2976368	species
- Diaphonia dispar	2976369	species
- Diaphonia luteola	2740248	species
- Diaphonia xanthopyga	2740251	species
unclassified Diaphonia	2740249	no rank
Dicronocephalus	1207178	genus
Dicronocephalus adamsi	1207179	species
- Dicronocephalus dabryi	1573400	species
Dicronocephalus uenoi	1573396	species
Dicronocephalus wallichii	1573397	species
Dicronocephalus yui	1573398	species
Dicronorhina	2900061	genus
- Dicronorhina derbyana	2900062	species
Dilochrosis	1207180	genus
- Dilochrosis atripennis	2740204	species
- Dilochrosis balteata	1207181	species
- Dilochrosis brownii	2740205	species
- Dilochrosis walteri	2740206	species
unclassified Dilochrosis	2619978	no rank
Diploa	1208524	genus
unclassified Diploa	2636841	no rank
Diplognatha	1207138	genus
- Diplognatha blanchardi	1207139	species
- Diplognatha cf. gagates DA-2012	1207326	species
- Diplognatha gagates	1207182	species
unclassified Diplognatha	2630524	no rank
Dischista	1207183	genus
- Dischista cincta	1207184	species
Elaphinis	1207140	genus
- Elaphinis delagoensis	1207141	species
- Elaphinis irrorata	1207142	species
Eudicella	1207145	genus
- Eudicella darwiniana	1678847	species
- Eudicella gralli	1678848	species
- Eudicella morgani	1678849	species
- Eudicella nana	1678850	species
- Eudicella pauperata	1678851	species
- Eudicella quadrimaculata	2900050	species
Eudicella schultzeorum	1207146	species
- Eudicella smithii	2900051	species
Eudicella tetraspilota	2900052	species
- Eudicella trilineata	1207148	species
Euphoria	1541327	genus
- Euphoria fulgida	2680441	species
- Euphoria kernii	2680442	species
- Euphoria sepulcralis	2680443	species
- Euphoria verticalis	2680444	species
unclassified Euphoria	2634909	no rank
Eupoecila	1552012	genus
- Eupoecila australasiae	2740207	species
- Eupoecila evanescens	2740208	species
- Eupoecila inscripta	2740209	species
unclassified Eupoecila	2630317	no rank
Euryomia	1207185	genus
- Euryomia argentea	1207186	species
Euselates	1207187	genus
- Euselates antennata	1207188	species
unclassified Euselates	2632197	no rank
Evanides	2740252	genus
- Evanides rufolatera	2740253	species
Gametis	1310344	genus
- Gametis bealiae	1618288	species
- Gametis jucunda	1310345	species
- Gametis versicolor	1499646	species
unclassified Gametis	3069454	no rank
Glycyphana	1207149	genus
- Glycyphana binotata	1207150	species
- Glycyphana brunnipes	2740210	species
- Glycyphana fulvistemma	1885608	species
- Glycyphana horsfieldi	1207151	species
- Glycyphana nicobarica	1618285	species
- Glycyphana papua	2740211	species
- Glycyphana pulchra	2740212	species
- Glycyphana stolata	2680496	species
unclassified Glycyphana	2624540	no rank
Gnorimella	1932083	genus
- Gnorimella maculosa	1932084	species
Goliathus	1207152	genus
- Goliathus goliatus	1207153	species
- Goliathus orientalis	1207189	species
Hemichnoodes	2740254	genus
- Hemichnoodes mniszechi	2740255	species
- Hemichnoodes parryi	2740256	species
Hemilia	1207190	genus
- Hemilia striata	1207191	species
Hemipharis	1207192	genus
- Hemipharis insularis	1207193	species
unclassified Hemipharis	2621967	no rank
Heterocnemis	465252	genus
- Heterocnemis graeca	465253	species
Heterorrhina	383512	genus
- Heterorrhina cf. sexmaculata DA-2012	1207330	species
- Heterorrhina elegans	1388893	species
- Heterorrhina micans	383522	species
unclassified Heterorrhina	2647616	no rank
Inca	1207154	genus
Inca clathrata	1207155	species
Ischiopsopha	1552026	genus
- Ischiopsopha pulchripes	2740213	species
- Ischiopsopha wallacei	2740214	species
unclassified Ischiopsopha	2629776	no rank
Ixorida	3394797	genus
- Ixorida albonotata	3394798	species
Jumnos	1207194	genus
Jumnos ruckeri	1207195	species
Laevocelis	207276	genus
unclassified Laevocelis	2643319	no rank
Lasiotrichius	1618310	genus
- Lasiotrichius succinctus	1618311	species
Lenosoma	2740257	genus
- Lenosoma fulgens	2740258	species
Leucocelis	302191	genus
- Leucocelis amethystina	1207080	species
- Leucocelis haemorrhoidalis	1207089	species
- Leucocelis rubra	1207090	species
unclassified Leucocelis	2633919	no rank
Lomaptera	1552030	genus
- Lomaptera australis	2740215	species
- Lomaptera cinnamomea	2740216	species
unclassified Lomaptera	2640353	no rank
Lyraphora	1552034	genus
- Lyraphora obliquata	2740217	species
- Lyraphora velutina	2740218	species
unclassified Lyraphora	2621259	no rank
Mausoleopsis	1207196	genus
- Mausoleopsis amabilis	1207197	species
Mecynorhina	2592932	genus
- Mecynorhina polyphemus	2900054	species
Mecynorhina torquata	2592933	species
Melinesthes	1207217	genus
- Melinesthes algoensis	1207218	species
unclassified Melinesthes	2677637	no rank
Meroloba	1207219	genus
- Meroloba suturalis	1207220	species
Metallesthes	2740259	genus
- Metallesthes anneliesae	2740260	species
- Metallesthes metallescens	2740261	species
- Metallesthes unicolor	2740264	species
unclassified Metallesthes	2740262	no rank
Micropoecila	2740265	genus
- Micropoecila cincta	2740266	species
Microvalgus	465291	genus
- Microvalgus glaber	2740219	species
- Microvalgus nigrinus	2740220	species
- Microvalgus vagans	2740222	species
unclassified Microvalgus	2638360	no rank
Mycterophallus	1552044	genus
- Mycterophallus duboulayi	2740223	species
unclassified Mycterophallus	2635372	no rank
Myodermum	1205635	genus
unclassified Myodermum	2627503	no rank
Navigator	2740267	genus
- Navigator fossor	2740268	species
- Navigator pixii	2740269	species
Neoclithria	2740270	genus
- Neoclithria eburneoguttata	2740271	species
Neorrhina	1552052	genus
- Neorrhina octopunctata	2740224	species
- Neorrhina punctata	2740225	species
unclassified Neorrhina	2629469	no rank
Neptunides	1207221	genus
- Neptunides stanleyi	1207222	species
Niphetophora	1208528	genus
unclassified Niphetophora	2649336	no rank
Nipponovalgus	1885629	genus
- Nipponovalgus angusticollis	1885630	species
Octocollis	2740272	genus
- Octocollis setosus	2740273	species
Oplostomus	1230912	genus
- Oplostomus fuligineus	1232491	species	large hive beetle
- Oplostomus haroldi	1230913	species
Osmoderma	174903	genus
- Osmoderma barnabita	314303	species
- Osmoderma caeleste	1940696	species
- Osmoderma cristinae	314301	species
- Osmoderma eremicola	1932075	species
- Osmoderma eremita	314302	species
- Osmoderma italicum	314305	species
- Osmoderma lassallei	314304	species
- Osmoderma opicum	1884344	species
unclassified Osmoderma	2627208	no rank
Oxythyrea	465316	genus
- Oxythyrea albopicta	2974024	species
- Oxythyrea cinctella	465317	species
- Oxythyrea dulcis	2974025	species
- Oxythyrea funesta	1207102	species
- Oxythyrea pantherina	1207103	species
unclassified Oxythyrea	2766503	no rank
Pachnoda	7057	genus
- Pachnoda cf. sinuata DA-2012	1207336	species
- Pachnoda fasciata	1207104	species
Pachnoda marginata	7058	species
- Pachnoda thoracica	1207107	species
unclassified Pachnoda	2612245	no rank
Paratrichius	1441448	genus
- Paratrichius doenitzi	1441449	species
Paratrichius duplicatus	1441453	species
- Paratrichius geibiensis	1441451	species
- Paratrichius itoi	1441450	species
- Paratrichius kyushuensis	1441452	species
- Paratrichius nomurai	1441455	species
Phoxomela	1207223	genus
- Phoxomela umbrosa	1207224	species
Phoxomeloides	1230910	genus
- Phoxomeloides gedyei	1230911	species
Phyllopodium	2740287	genus
- Phyllopodium palmatum	2740288	species
Plaesiorrhinella	1207225	genus
- Plaesiorrhinella plana	1207226	species
- Plaesiorrhinella trivittata	1207227	species
Plectrone	1207228	genus
Plectrone borneensis	1207229	species
Poecilopharis	2740274	genus
- Poecilopharis leai	2740275	species
Polybaphes	1207231	genus
- Polybaphes balteata	1207232	species
Porphyronota	1207233	genus
Porphyronota hebraea	1207234	species
- Porphyronota maculatissima	1207236	species
Protaetia	348687	genus
Calopotosia	2100446	subgenus
Cetonischema	2100447	subgenus
Eumimimetica	2792003	subgenus
Eupotosia	2100442	subgenus
Indoprotaetia	2100443	subgenus
Liocola	2100445	subgenus
Netocia	1208526	subgenus
Philhelena	2100448	subgenus
Potosia	1208530	subgenus
Potosiomima	2100444	subgenus
Protaetia incertae sedis	2100450	no rank
Protaetia speciosa	2900056	species
unclassified Protaetia	2100449	no rank
Pseudinca	1207198	genus
- Pseudinca marmoratus	1207199	species
Pseudochalcothea	1207157	genus
- Pseudochalcothea spathulifera	1207158	species
Pseudoclithria	1552072	genus
- Pseudoclithria adusta	2740226	species
- Pseudoclithria anchoralis	2740227	species
- Pseudoclithria dejecta	2740228	species
- Pseudoclithria hirticeps	2740229	species
unclassified Pseudoclithria	2626912	no rank
Pseudotorynorrhina	2859704	genus
- Pseudotorynorrhina japonica	2859705	species
Rhabdotis	1207200	genus
- Rhabdotis aulica	1207201	species
Rhomborrhina	1163332	genus
- Rhomborrhina japonica	1163331	species
Scaptobius	1208532	genus
unclassified Scaptobius	2628520	no rank
Schizorhina	2740276	genus
- Schizorhina atropunctata	2740277	species
- Schizorhina immaculata	2740278	species
Stenopisthes	2740279	genus
- Stenopisthes frenchi	2740280	species
Stephanorrhina	1207159	genus
- Stephanorrhina princeps	1207160	species
Storeyus	2740281	genus
- Storeyus fasciculatum	2740282	species
Stripsipher	1142449	genus
- Stripsipher zebra	1207092	species
unclassified Stripsipher	2622348	no rank
Taeniodera	1207161	genus
Taeniodera zebraea	1207162	species
unclassified Taeniodera	2631388	no rank
Tapinoschema	2740283	genus
- Tapinoschema digglesii	2740284	species
- Tapinoschema impar	2740285	species
- Tapinoschema lacunosa	2740286	species
Tephraea	1207205	genus
- Tephraea cinerea	1564139	species
- Tephraea dichroa	1207206	species
- Tephraea morosa	1207207	species
Thaumastopeus	1207208	genus
- Thaumastopeus pugnator	1207209	species
- Thaumastopeus shangaicus	1453209	species
Trichaulax	1552119	genus
- Trichaulax kirbyi	2740232	species
- Trichaulax marginipennis	2740233	species
- Trichaulax philipsii	2740234	species
unclassified Trichaulax (in: beetles)	2689016	no rank
Trichiotinus	1710621	genus
- Trichiotinus affinis	1932076	species
- Trichiotinus assimilis	1710622	species
- Trichiotinus bibens	1932077	species
- Trichiotinus lunulatus	1932078	species
- Trichiotinus piger	1932079	species
- Trichiotinus rufobrunneus	1932080	species
- Trichiotinus texanus	1932081	species
- Trichiotinus viridans	1932082	species
Trichius	465339	genus
- Trichius fasciatus	1207093	species
- Trichius rosaceus	1207094	species
- Trichius sexualis	1588518	species
- Trichius succinctus	1310326	species
- Trichius zonatus	465340	species
unclassified Trichius	2629838	no rank
Trichostetha	1207210	genus
- Trichostetha fascicularis	1207211	species
Trigonopeltastes	1932085	genus
- Trigonopeltastes delta	1932086	species
Trigonophorus	1822357	genus
unclassified Trigonophorus	2639611	no rank
Tropinota	980915	genus
- Tropinota hirta	1207095	species
- Tropinota squalida	980916	species
Xeloma	1207212	genus
- Xeloma leprosa	1207213	species
- Xeloma tomentosa	1207214	species
unclassified Cetoniinae	1207110	no rank
- Cetoniinae sp. 'Bombila' DA-2012	1339110	species
- Cetoniinae sp. NLG-2014	1551876	species
- Crematoscheilina sp. DA-2012	1208514	species
- Taenioderini sp. 1 DA-2012	1207345	species
- Taenioderini sp. 2 DA-2012	1207346	species
Dynamopodinae	3085250	subfamily
Orubesa	3085251	genus
- Orubesa ata	3085252	species
- Orubesa plicifrons	3085253	species
Dynastinae	50519	subfamily	rhinoceros beetles, hercules beetles & elephant beetles
Adoryphorus	1069820	genus
- Adoryphorus coulonii	1069821	species
Aegopsis	1411910	genus
- Aegopsis curvicornis	2728559	species
Agaocephala	1247150	genus
- Agaocephala cornigera	1247151	species
Alissonotum	383509	genus
- Alissonotum binodulum	383518	species
- Alissonotum simile	383519	species
unclassified Alissonotum	3069445	no rank
Allomyrina	50412	genus
- Allomyrina davidis	1248198	species
- Allomyrina pfeifferi	1247134	species
Amblyoproctus	1261324	genus
- Amblyoproctus centroamericanus	1261325	species
Ancognatha	1247152	genus
- Ancognatha gracilis	2728561	species
- Ancognatha vexans	1261269	species
- Ancognatha vulgaris	1247153	species
unclassified Ancognatha	2728628	no rank
Aspidolea	1261326	genus
- Aspidolea fuliginea	2728565	species
- Aspidolea singularis	1261327	species
unclassified Aspidolea	2728635	no rank
Augosoma	1247154	genus
- Augosoma centaurus	1247155	species
Beckius	1247141	genus
- Beckius beccarii	1247142	species
Bothynus	1247177	genus
- Bothynus quadridens	2728567	species
unclassified Bothynus	2621246	no rank
Calicnemis	1496342	genus
- Calicnemis latreillii	1496343	species
- Calicnemis obesa	1496344	species
- Calicnemis sardiniensis	1496345	species
Carneodon	1552410	genus
unclassified Carneodon	2634034	no rank
Chalcosoma	1247143	genus
Chalcosoma atlas	1247156	species
Chalcosoma caucasus	1247144	species
- Chalcosoma chiron	1247145	species
Cheiroplatys	465195	genus
- Cheiroplatys latipes	465196	species
unclassified Cheiroplatys	2677132	no rank
Coelosis	1274983	genus
- Coelosis biloba	1274984	species
Cryptodus	1551999	genus
unclassified Cryptodus	2645192	no rank
Cyclocephala	465218	genus
- Cyclocephala aequatoria	1714552	species
- Cyclocephala aff. pasadenae/hirta MV20140605	2116607	species
- Cyclocephala aff. pasadenae/hirta MV20150618	2116608	species
- Cyclocephala aff. pasadenae/hirta MV20151129	2116609	species
- Cyclocephala almitana	1261277	species
- Cyclocephala amazona	1714553	species
- Cyclocephala amblyopsis	1261278	species
- Cyclocephala atripes	1940448	species
- Cyclocephala borealis	1546013	species	northern masked chafer
- Cyclocephala brevis	1261279	species
- Cyclocephala brittoni	2728572	species
- Cyclocephala carbonaria	1261280	species
- Cyclocephala concolor	1274965	species
- Cyclocephala confusa	1261281	species
- Cyclocephala conspicua	1261282	species
- Cyclocephala deceptor	2591315	species
- Cyclocephala discicollis	2728573	species
- Cyclocephala discolor	1714554	species
- Cyclocephala erotylina	1261283	species
- Cyclocephala fasciolata	1274966	species
- Cyclocephala freyi	2358228	species
- Cyclocephala gravis	1261284	species
- Cyclocephala inca	1714555	species
- Cyclocephala insulicola	1133717	species
- Cyclocephala kaszabi	1261285	species
- Cyclocephala krombeini	2728574	species
- Cyclocephala ligyrina	1261286	species
- Cyclocephala longula	2093234	species
- Cyclocephala lunulata	1261287	species
- Cyclocephala lurida	2680285	species
Cyclocephala mafaffa	1133718	species
Cyclocephala melanocephala	1133720	species
- Cyclocephala multiplex	1714556	species
- Cyclocephala nigerrima	1261288	species
- Cyclocephala nigritarsis	1261289	species
- Cyclocephala pan	1274967	species
- Cyclocephala porioni	1261290	species
- Cyclocephala quadripunctata	2728575	species
- Cyclocephala sanguinicollis	1261291	species
- Cyclocephala sexpunctata	1274968	species
- Cyclocephala signaticollis	2680286	species
- Cyclocephala sparsa	1261292	species
- Cyclocephala stictica	1261293	species
- Cyclocephala stockwelli	1261294	species
Cyclocephala tridentata	1133722	species
- Cyclocephala tridentata tridentata x Cyclocephala insulicola	1133725	species
- Cyclocephala tutilina	1714557	species
- Cyclocephala weidneri	1261295	species
- Cyclocephala williami	1274969	species
- Cyclocephala zodion	1261296	species
unclassified Cyclocephala	2620195	no rank
Cyphonistes	1205659	genus
- Cyphonistes vallatus	1205660	species
Diloboderus	1718887	genus
- Diloboderus abderus	1718888	species
Dynastes	60842	genus
- Dynastes grantii	60843	species
Dynastes hercules	1247135	species
- Dynastes hyllus	1767015	species
- Dynastes maya	1767016	species
- Dynastes maya x Dynastes grantii	2950246	species
- Dynastes moroni	1767017	species
- Dynastes neptunus	1247136	species
- Dynastes satanas	1767018	species
- Dynastes tityus	1248199	species
Dyscinetus	1261339	genus
- Dyscinetus dubius	1261340	species
- Dyscinetus morator	1714558	species
Enema	1247158	genus
- Enema pan	1247159	species
Eophileurus	1247160	genus
Eophileurus chinensis	1247161	species
unclassified Eophileurus	2620563	no rank
Euetheola	2680597	genus
- Euetheola humilis	2680598	species
- Euetheola rugiceps	3120940	species
Eupatorus	1247163	genus
- Eupatorus birmanicus	1248200	species
- Eupatorus gracilicornis	1247164	species
- Eupatorus hardwickei	2950138	species
- Eupatorus sukkiti	2944873	species
Eutyctus	1496346	genus
- Eutyctus deserti	1496347	species
Golofa	1247146	genus
- Golofa clavigera	1248197	species
- Golofa eacus	1247165	species
- Golofa pizarro	1248201	species
- Golofa porteri	1247147	species
Haploscapanes	1248208	genus
- Haploscapanes barbarossa	1248209	species
Hemiphileurus	1261341	genus
- Hemiphileurus dyscritus	1261342	species
- Hemiphileurus pygidiopunctissimus	2728585	species
- Hemiphileurus variolosus	1261343	species
Heterogomphus	1261344	genus
- Heterogomphus mniszechi	1261345	species
Hexodon	1247166	genus
- Hexodon unicolor	1247167	species
Homophileurus	1261346	genus
- Homophileurus quadrituberculatus	1261347	species
- Homophileurus tricuspis	2728587	species
Lycomedes	1247168	genus
- Lycomedes buckleyi	1247169	species
Marronus	2654887	genus
- Marronus borbonicus	2654888	species
Megaceras	1261348	genus
- Megaceras septentrionis	1261349	species
Megasoma	575205	genus
- Megasoma actaeon	1247137	species
Megasoma elephas	575206	species
- Megasoma mars	1247138	species
- Megasoma thersites	1248203	species
Mellissius	3032549	genus
- Mellissius popei	3032550	species
Metanastes	1552038	genus
unclassified Metanastes	2632081	no rank
Mimeoma	1714559	genus
- Mimeoma acuta	1714560	species
- Mimeoma englemani	1714561	species
- Mimeoma maculata	1714562	species
- Mimeoma nigra	1714563	species
- Mimeoma signatoides	1714564	species
Neocorynophyllus	1552046	genus
unclassified Neocorynophyllus	2628466	no rank
Novapus	1552058	genus
unclassified Novapus	2630347	no rank
Orizabus	1248214	genus
- Orizabus clunalis	1248215	species
- Orizabus pyriformis	2680762	species
Oryctes	72549	genus
- Oryctes agamemnon	915460	species
- Oryctes borbonicus	1629725	species
- Oryctes cf. boas DA-2012	1207333	species
- Oryctes chevrolatii	2709812	species
- Oryctes elegans	2213224	species
- Oryctes gnu	1247139	species
- Oryctes mayottensis	2709813	species
- Oryctes minor	2709814	species
Oryctes nasicornis	465150	species
- Oryctes rhinoceros	72550	species	rhinoceros beetle
- Oryctes tarandus	2709815	species
unclassified Oryctes (in: beetles)	2612243	no rank
Oryctoderus	1247170	genus
- Oryctoderus latitarsis	1247171	species
unclassified Oryctoderus	2634890	no rank
Pachyoryctes	1248210	genus
- Pachyoryctes solidus	1248211	species
Papuana	1247172	genus
Papuana woodlarkiana	1247173	species
Parapucaya	1261350	genus
- Parapucaya amazonica	1261351	species
Peltonotus	1542271	genus
- Peltonotus nasutus	1542272	species
Pentodon	207356	genus
- Pentodon algerinum	207357	species
- Pentodon bidens	1588352	species
- Pentodon idiota	465151	species
Pentodon quadridens	1618289	species
unclassified Pentodon (in: beetles)	2688796	no rank
Pericoptus	3375621	genus
unclassified Pericoptus	3375622	no rank
Phileurus	1247187	genus
- Phileurus aff. didymus AS1	2728998	species
- Phileurus carinatus	2728601	species
- Phileurus didymus	1261301	species
- Phileurus limicauda	2728602	species
- Phileurus truncatus	1274971	species
- Phileurus valgus	2728603	species
- Phileurus youngi	1261302	species
- Phileurus zonianis	2728604	species
unclassified Phileurus	2635533	no rank
Phyllognathus	465330	genus
- Phyllognathus dionysius	465331	species
- Phyllognathus excavatus	2599481	species
unclassified Phyllognathus	3069463	no rank
Pimelopus	465332	genus
Pimelopus dubius	465333	species
Podischnus	2728700	genus
- Podischnus agenor	2728701	species
Pucaya	1261356	genus
- Pucaya castanea	1261357	species
- Pucaya pulchra	2358229	species
Pycnoschema	1208536	genus
unclassified Pycnoschema	2646863	no rank
Semanopterus	1552089	genus
unclassified Semanopterus	2626233	no rank
Spodistes	1261358	genus
- Spodistes batesi	2728615	species
- Spodistes beltianus	1261359	species
Stenocrates	1261362	genus
- Stenocrates laevicollis	1261363	species
Strategus	984468	genus
- Strategus aff. aloeus AS2	2728999	species
- Strategus aloeus	984469	species
- Strategus jugurtha	2728617	species
Syrictes	1207202	genus
- Syrictes syrichthus	1207203	species
Temnorhynchus	1208534	genus
unclassified Temnorhynchus	2618861	no rank
Tomarus	1261364	genus
- Tomarus bituberculatus	2728623	species
- Tomarus gibbosus	1710662	species
- Tomarus sallaei	1261365	species
unclassified Tomarus	3031270	no rank
Trichogomphus	1247174	genus
- Trichogomphus martabani	1247175	species
- Trichogomphus mongol	2910522	species
Trissodon	1552104	genus
unclassified Trissodon	2634877	no rank
Trypoxylus	273927	genus
Trypoxylus dichotomus	273928	species
- Trypoxylus kanamorii	2710795	species
unclassified Trypoxylus	3069087	no rank
Xyloryctes	50520	genus
- Xyloryctes faunus	50521	species
- Xyloryctes gideon	207358	species
Xylotrupes	1247148	genus
Xylotrupes australicus	1248205	species
Xylotrupes beckeri	1247149	species
- Xylotrupes carinulus	2172590	species
- Xylotrupes clinias	2172591	species
- Xylotrupes damarensis	2172592	species
- Xylotrupes falcatus	2172593	species
Xylotrupes florensis	2172594	species
- Xylotrupes gideon	1247176	species
- Xylotrupes inarmatus	2172596	species
- Xylotrupes lorquini	2172597	species
Xylotrupes macleayi	2172598	species
- Xylotrupes meridionalis	1248206	species
Xylotrupes mniszechi	1689722	species
- Xylotrupes pachycera	2172600	species
- Xylotrupes pauliani	2172601	species
Xylotrupes philippinensis	2172602	species
Xylotrupes pubescens	2172605	species
- Xylotrupes rindaae	2172607	species
Xylotrupes socrates	2900059	species
Xylotrupes sumatrensis	2172608	species
- Xylotrupes tadoana	2172610	species
- Xylotrupes telemachos	2172611	species
- Xylotrupes ulysses	2172612	species
- Xylotrupes wiltrudae	2172613	species
unclassified Xylotrupes	2685637	no rank
unclassified Dynastinae	207281	no rank
- Cyclocephalini sp. Modo	1247180	species
- Dynastinae sp. BOLD:AAG5711	2658847	species
- Dynastinae sp. DA-2012	1207340	species
- Dynastinae sp. NLG-2014	1551885	species
- Pentodontini sp. Kuro	1247186	species
- Phileurini sp. PHILoz	207282	species
Melolonthinae	7059	subfamily	chafers; May beetles & June bugs
Ablabera	1237320	genus
unclassified Ablabera	2631329	no rank
Achloa	1208538	genus
unclassified Achloa	2634108	no rank
Acylochilus	702311	genus
unclassified Acylochilus	2642349	no rank
Amblonoxia	1355133	genus
- Amblonoxia fieldi	2680088	species
- Amblonoxia palpalis	1355134	species
Amorphochelus	980917	genus
- Amorphochelus retusus	1629724	species
unclassified Amorphochelus	2647903	no rank
Amphimallon	221091	genus
- Amphimallon majale	1720622	species	European chafer
- Amphimallon solstitiale	360071	species
unclassified Amphimallon	3058215	no rank
Ancylonyx	1551964	genus
unclassified Ancylonyx	2637968	no rank
Antitrogus	247270	genus
- Antitrogus consanguineus	272057	species
- Antitrogus parvulus	247271	species	childers canegrub
- Antitrogus rugulosus	272056	species
Apiencya	465172	genus
unclassified Apiencya	2677005	no rank
Aplidia	1142435	genus
- Aplidia transversa	1142436	species
Apogonia	207354	genus
- Apogonia cf. basalis	2962665	species
- Apogonia comosa	3069448	species
- Apogonia coriacea	3069449	species
- Apogonia cribricollis	761910	species
- Apogonia cupreoviridis	761911	species
- Apogonia kombirana	3394783	species
- Apogonia nietneri	3069451	species
- Apogonia rauca	3379181	species
- Apogonia solida	3069450	species
- Apogonia splendida	2962664	species
unclassified Apogonia	2618348	no rank
Apomorphochelus	465174	genus
unclassified Apomorphochelus	2642706	no rank
Arctodium	2570514	genus
- Arctodium vulpinum	2570515	species
Asactopholis	3087141	genus
- Asactopholis dalatensis	3087142	species
Astenopholis	1237325	genus
unclassified Astenopholis	2624326	no rank
Asthenopholis	1205810	genus
unclassified Asthenopholis	2630523	no rank
Aulacoserica	1237327	genus
unclassified Aulacoserica	2685619	no rank
Automolius	1551974	genus
- Automolius humilis	465184	species
unclassified Automolius	2618568	no rank
Autoserica	1237351	genus
- Autoserica punctulata	1705045	species
- Autoserica tessellata	1237352	species
- Autoserica zoutpaniana	1698073	species
Balbera	1695848	genus
unclassified Balbera	2637405	no rank
Bilga	1695850	genus
unclassified Bilga	2627782	no rank
Blepharotoma	2904872	genus
unclassified Blepharotoma	2904873	no rank
Brahmina	1269194	genus
- Brahmina coriacea	1475332	species
- Brahmina crenicollis	1290864	species
- Brahmina faldermanni	1290865	species
- Brahmina mysoreensis	2704556	species
- Brahmina sedakovi	1618283	species
unclassified Brahmina	3069452	no rank
Brenskiella	1207121	genus
- Brenskiella flavomicans	1207122	species
Buettikeria	465187	genus
- Buettikeria echinosa	465188	species
Camenta	1237329	genus
- Camenta innocua	1661818	species
unclassified Camenta	2639066	no rank
Camentoserica	1237332	genus
- Camentoserica cf. livida DA-2012	1237333	species
- Camentoserica livida	1237353	species
Ceratochelus	980921	genus
unclassified Ceratochelus	2629141	no rank
Ceratogonia	465189	genus
- Ceratogonia bicornuta	465190	species
Chariochilus	1551986	genus
unclassified Chariochilus	2628784	no rank
Chasmatopterus	465191	genus
- Chasmatopterus villosus	465192	species
unclassified Chasmatopterus	2635365	no rank
Cheirotonus	1207172	genus
- Cheirotonus formosanus	3095916	species
- Cheirotonus gestroi	1207173	species
- Cheirotonus jansoni	1297118	species
Cherbezatina	465197	genus
- Cherbezatina strigosa	465198	species
Clania	1033757	genus
unclassified Clania (in: beetles)	2688349	no rank
Colobostoma	1551995	genus
unclassified Colobostoma	2633793	no rank
Colpochila	465203	genus
unclassified Colpochila	2643150	no rank
Colymbomorpha	1551997	genus
unclassified Colymbomorpha	2621940	no rank
Congella	980924	genus
- Congella cf. valida DA-2011	980926	species
unclassified Congella	2627551	no rank
Costelytra	50578	genus
- Costelytra brunnea	1090973	species
- Costelytra zealandica	50579	species
Cylinchus	980927	genus
- Cylinchus cf. 'vanus' DA-2011	1033752	species
unclassified Cylinchus	2634641	no rank
Cyphochilus	1453292	genus
Cyphochilus crataceus	1453293	species
- Cyphochilus podicalis	3087053	species
- Cyphochilus tonkinensis	3087054	species
- Cyphochilus ventritectus	3087055	species
Cyphonoxia	207352	genus
- Cyphonoxia kircheri	207353	species
Dasylepida	454918	genus
- Dasylepida ishigakiensis	454919	species	white grub beetle
Dedalopterus	2563635	genus
- Dedalopterus malyszi	3087057	species
Dermolepida	1552003	genus
- Dermolepida albohirtum	1980614	species
unclassified Dermolepida	2625780	no rank
Dichecephala	465222	genus
- Dichecephala ovata	465223	species
Dichelomorpha	3383716	genus
- Dichelomorpha alsiosia	3383717	species
Dichelonyx	1710458	genus
- Dichelonyx albicollis	1710459	species
- Dichelonyx backii	2339858	species
- Dichelonyx kirbyi	1845034	species
- Dichelonyx linearis	1710460	species
- Dichelonyx subvittata	1710461	species
- Dichelonyx sulcata	2680348	species
unclassified Dichelonyx	2636457	no rank
Dikellites	1552005	genus
unclassified Dikellites	2648095	no rank
Dinacoma	2592785	genus
- Dinacoma caseyi	2592786	species
- Dinacoma marginata	2592787	species
Diphucephala	1542303	genus
unclassified Diphucephala	2635549	no rank
Diphycerus	383511	genus
unclassified Diphycerus	2640243	no rank
Diplotaxis	1710485	genus
- Diplotaxis belfragei	2680367	species
- Diplotaxis brevicollis	1710486	species
- Diplotaxis frondicola	2680368	species
- Diplotaxis haydenii	1710487	species
- Diplotaxis liberta	2831976	species
- Diplotaxis mentalis	2680369	species
- Diplotaxis obscura	2109411	species
- Diplotaxis subangulata	2680370	species
- Diplotaxis tristis	1710488	species
unclassified Diplotaxis (in: beetles)	2658736	no rank
Doleroserica	1695852	genus
unclassified Doleroserica	2618243	no rank
Dolerotarsa	1237334	genus
- Dolerotarsa cf. emendatrix DA-2012	1237335	species
- Dolerotarsa emendatrix	1237354	species
unclassified Dolerotarsa	2632177	no rank
Dolichoplia	1033760	genus
unclassified Dolichoplia	2640352	no rank
Echyra	465226	genus
- Echyra umbrina	465227	species
Ectinohoplia	1630947	genus
- Ectinohoplia obducta	2682834	species
- Ectinohoplia rufipes	1630948	species
unclassified Ectinohoplia	2630607	no rank
Empecamenta	1237337	genus
unclassified Empecamenta	2630577	no rank
Empecta	465228	genus
- Empecta sicardi	465229	species
Enaria	465230	genus
- Enaria boissayei	465231	species
Epholcis	1552007	genus
unclassified Epholcis	2640691	no rank
Eriesthis	980898	genus
- Eriesthis cf. 'rhodesiana' DA-2011	1033754	species
- Eriesthis cf. 'schoenherri' DA-2011	1033753	species
- Eriesthis hypocrita	980899	species
- Eriesthis semihirta	980900	species
- Eriesthis vestita	980901	species
- Eriesthis vulpina	980902	species
Etiserica	1237339	genus
unclassified Etiserica	2620890	no rank
Euchirus	1207143	genus
- Euchirus dupontianus	1207144	species
- Euchirus longimanus	1968892	species
Euphoresia	1695854	genus
unclassified Euphoresia	2642828	no rank
Euthora	465239	genus
unclassified Euthora (in: beetles)	2677355	no rank
Eutrichesis	465241	genus
- Eutrichesis pilosicollis	465242	species
Geniates	980929	genus
unclassified Geniates	2620676	no rank
Geotrogus	3139948	genus
- Geotrogus genei	3139949	species
Gnaphalopoda	1552016	genus
unclassified Gnaphalopoda	2645853	no rank
Gymnoloma	1033762	genus
unclassified Gymnoloma	2626724	no rank
Hadrops	1552020	genus
unclassified Hadrops	2622196	no rank
Haplidia	465247	genus
- Haplidia cypria	3229163	species
Haplidia transversa	465248	species
unclassified Haplidia	2622398	no rank
Harpechys	1552022	genus
unclassified Harpechys	2648868	no rank
Hemiserica	397611	genus
- Hemiserica nasutella	397612	species
Heptophylla	91460	genus
Heptophylla picea	91461	species	yellowish elongate chafer
Heterochelus	980903	genus
- Heterochelus cf. 'lydenburgensis' DA-2011	1033755	species
- Heterochelus defector	980904	species
- Heterochelus sulphureus	980905	species
- Heterochelus vulpinus	980906	species
unclassified Heterochelus	2621450	no rank
Heteronyx	365801	genus
- Heteronyx pubescens	1524718	species
unclassified Heteronyx	2685854	no rank
Hilyotrogus	1290874	genus
- Hilyotrogus bicoloreus	1290875	species
Holochelus	1588542	genus
- Holochelus vernus	1588543	species
Holotrichia	33393	genus
- Holotrichia consanguinea	1475333	species
- Holotrichia diomphalia	33394	species
- Holotrichia ernesti	1618298	species
- Holotrichia fissa	1916670	species
- Holotrichia longipennis	1475334	species
- Holotrichia nagpurensis	1475335	species
- Holotrichia niponensis	2872680	species
- Holotrichia oblita	644536	species
- Holotrichia parallela	93412	species	large black chafer
- Holotrichia plumbea	879211	species
- Holotrichia reynaudi	1916671	species
- Holotrichia rufoflava	2704557	species
- Holotrichia serrata	1032692	species
- Holotrichia seticollis	383502	species
- Holotrichia sikkimensis	1475336	species
unclassified Holotrichia	2638877	no rank
Homalochilus	2904695	genus
- Homalochilus punctatostriatus	2904696	species
Homaloplia	207336	genus
unclassified Homaloplia	2642734	no rank
Homalotropus	465254	genus
unclassified Homalotropus	2635481	no rank
Homolotropus	1552024	genus
unclassified Homolotropus	2637064	no rank
Hoplia	190493	genus
- Hoplia africana	190496	species
- Hoplia argentea	190498	species
- Hoplia bilineata	190497	species
- Hoplia chlorophana	190494	species
- Hoplia cincticollis	1822361	species
- Hoplia farinosa	980884	species
- Hoplia freyi	190495	species
- Hoplia graminicola	465125	species
- Hoplia korbi	980885	species
Hoplia philanthus	190499	species
- Hoplia praticola	2561399	species
- Hoplia trifasciata	2486041	species
unclassified Hoplia	2641918	no rank
Hoplochelus	465256	genus
- Hoplochelus marginalis	1629726	species
- Hoplochelus piliger	465257	species
Hoplosternus	1290876	genus
- Hoplosternus incanus	1290877	species
Hybocamenta	1695861	genus
unclassified Hybocamenta	2624613	no rank
Hyboserica	1237344	genus
unclassified Hyboserica	2646339	no rank
Hypopholis	1208540	genus
- Hypopholis cf. sommeri DA-2012	1237209	species
unclassified Hypopholis	2624345	no rank
Idaecamenta	1695863	genus
unclassified Idaecamenta	2643528	no rank
Idionychus	383514	genus
- Idionychus excisus	383523	species
Isonychus	465270	genus
- Isonychus pictus	1274970	species
unclassified Isonychus	2622336	no rank
Lepidiota	207350	genus
- Lepidiota albistigma	383503	species
- Lepidiota frenchi	272053	species
- Lepidiota mansueta	1188230	species
- Lepidiota nana	207351	species
- Lepidiota negatoria	272055	species
- Lepidiota noxia	272054	species
- Lepidiota sticticoptera	3291024	species
- Lepidiota stradbrokensis	465128	species
unclassified Lepidiota	2642071	no rank
Lepidoserica	1695867	genus
unclassified Lepidoserica	2623395	no rank
Lepithrix	1033766	genus
- Lepithrix cf. 'forsteri' DA-2011	1033767	species
Leucopholis	1075168	genus
- Leucopholis burmeisteri	1075169	species
- Leucopholis coneophora	2283329	species
- Leucopholis lepidophora	1902984	species
unclassified Leucopholis	3069456	no rank
Lichnia	1542247	genus
- Lichnia limbata	1542248	species
Liogenys	980936	genus
- Liogenys flaveola	2978517	species
- Liogenys grandis	2904688	species
- Liogenys hirta	2904689	species
- Liogenys latitarsis	2904690	species
- Liogenys mendozana	2904691	species
- Liogenys obesula	2978518	species
- Liogenys obscura	2904692	species
- Liogenys palpalis	2904693	species
- Liogenys wagenknechti	2904694	species
unclassified Liogenys	2637228	no rank
Liparetrus	465281	genus
- Liparetrus obscurior	1143062	species
unclassified Liparetrus	2620396	no rank
Lobogeniates	980938	genus
unclassified Lobogeniates	2623178	no rank
Macrodactylus	465283	genus
- Macrodactylus cf. subspinosus	3385162	species
- Macrodactylus subspinosus	2680668	species
unclassified Macrodactylus	2642849	no rank
Maechidius	465286	genus
unclassified Maechidius	2644633	no rank
Melolontha	7060	genus
- Melolontha cuprescens	2821240	species
- Melolontha furcicauda	2821241	species
- Melolontha hippocastani	903833	species
- Melolontha indica	2821242	species
- Melolontha masafumii	1247140	species
- Melolontha melolontha	7061	species	maybug; cockchafer
- Melolontha pectoralis	903834	species
unclassified Melolontha	2623060	no rank
Mesoserica	1695870	genus
- Mesoserica transvaalensis	1704666	species
unclassified Mesoserica	2628312	no rank
Michaeloplia	465288	genus
- Michaeloplia montana	465289	species
- Michaeloplia prolata	465290	species
Microplus	980909	genus
- Microplus cf. nemoralis DA-2011	980940	species
Miridiba	2704561	genus
- Miridiba castanea	2915476	species
- Miridiba coromandeliana	2829784	species
- Miridiba excisa	2704562	species
- Miridiba parasinensis	3087096	species
- Miridiba saigonensis	3087097	species
- Miridiba trichophora	2986799	species
Monochelus	980911	genus
- Monochelus cf. jucundus spSA1	980941	species
unclassified Monochelus	2644928	no rank
Nedymoserica	1237356	genus
- Nedymoserica flavida	1237357	species
unclassified Nedymoserica	2620074	no rank
Neogutierrezia	702818	genus
- Neogutierrezia chelii	702819	species
- Neogutierrezia scutata	702820	species
Neoheteronyx	1552048	genus
unclassified Neoheteronyx	2647738	no rank
Neophyllotocus	1552050	genus
unclassified Neophyllotocus	2639149	no rank
Neso	1552054	genus
unclassified Neso	2619781	no rank
Nigrotrichia	2915477	genus
- Nigrotrichia convexopyga	2915479	species
- Nigrotrichia kiotonensis	2915480	species
Nigrotrichia loochooana	2915478	species
Nuiba	3087155	genus
- Nuiba portentosa	3087156	species
Odontria	50575	genus
- Odontria striata	50576	species
- Odontria variegata	50577	species
unclassified Odontria	2624283	no rank
Omocrates	980942	genus
unclassified Omocrates	2638207	no rank
Oocamenta	1695872	genus
unclassified Oocamenta	2677462	no rank
Ophropyx	1552062	genus
unclassified Ophropyx	2646225	no rank
Pachrodema	2904697	genus
- Pachrodema lucida	2904698	species
unclassified Pachrodema	2904869	no rank
Pachycnema	980913	genus
- Pachycnema calviniana	980914	species
Pachydema	614201	genus
- Pachydema menieri	3413686	species
unclassified Pachydema	2628380	no rank
Pachypus	465320	genus
- Pachypus caesus	702043	species
- Pachypus candidae	465321	species
- Pachypus demoflysi	3108251	species
- Pachypus melonii	3108252	species
- Pachypus sardiniensis	3108253	species
unclassified Pachypus	2641669	no rank
Pacuvia	2904699	genus
- Pacuvia castanea	2904700	species
Paramorphochelus	465324	genus
- Paramorphochelus agricola	465325	species
unclassified Paramorphochelus	2621048	no rank
Paronyx	1552066	genus
unclassified Paronyx	2630220	no rank
Pedinotrichia	2915483	genus
- Pedinotrichia picea	2915484	species
Periserica	2945121	genus
unclassified Periserica	2945122	no rank
Perissosoma	465328	genus
unclassified Perissosoma	2631742	no rank
Philoserica	1237347	genus
- Philoserica vittata	1237358	species
unclassified Philoserica	2677656	no rank
Phyllophaga	219349	genus
- Phyllophaga aemula	1890705	species
- Phyllophaga aequata	1261303	species
- Phyllophaga affabilis	489793	species
- Phyllophaga anxia	1710756	species
- Phyllophaga arkansana	1890706	species
- Phyllophaga balia	703313	species
- Phyllophaga batillifer	3248543	species
- Phyllophaga bipartita	489794	species
- Phyllophaga boruca	1274972	species
- Phyllophaga brevidens	1891723	species
- Phyllophaga calceata	489795	species
- Phyllophaga cf. fraterna 1 MPR-2016	1890722	species
- Phyllophaga cf. fraterna 2 MPR-2016	1890723	species
- Phyllophaga cf. fraterna 3 MPR-2016	1890724	species
- Phyllophaga changuena	1274973	species
- Phyllophaga chorotega	1261304	species
- Phyllophaga clypeata	1890707	species
- Phyllophaga congrua	489796	species
- Phyllophaga corrosa	489797	species
- Phyllophaga crassissima	489798	species
- Phyllophaga crenulata	489799	species
- Phyllophaga cribrosa	2680840	species
- Phyllophaga crinita	489800	species
- Phyllophaga cupuliformis	1890708	species
- Phyllophaga delata	703314	species
- Phyllophaga densata	1261305	species
- Phyllophaga dentex	3248544	species
- Phyllophaga diffinis	1890709	species
- Phyllophaga drakii	703315	species
- Phyllophaga ephilida	489801	species
- Phyllophaga errans	1890710	species
- Phyllophaga fervida	1890711	species
- Phyllophaga forsteri	703316	species
- Phyllophaga foxii	703317	species
Phyllophaga fraterna	703318	species
- Phyllophaga fusca	489802	species
- Phyllophaga futilis	489803	species
- Phyllophaga gigantea	1274974	species
- Phyllophaga glaberrima	1890712	species
- Phyllophaga glabricula	489804	species
- Phyllophaga guanacasteca	1274975	species
- Phyllophaga guapiles	1274976	species
- Phyllophaga guapiloides	1261306	species
- Phyllophaga hirsuta	703319	species
- Phyllophaga hirticula	703320	species
- Phyllophaga hirtiventris	489805	species
- Phyllophaga hornii	703321	species
- Phyllophaga ilicis	489806	species
- Phyllophaga implicita	489807	species
- Phyllophaga inepta	489808	species
- Phyllophaga inversa	1890713	species
- Phyllophaga knochii	703322	species
- Phyllophaga larai	1261307	species
- Phyllophaga latifrons	1890714	species
- Phyllophaga lenis	1891724	species
- Phyllophaga lissopyge	1261308	species
- Phyllophaga lodingi	703323	species
- Phyllophaga longispina	2486045	species
- Phyllophaga longitarsa	489809	species
- Phyllophaga lorencita	1261309	species
- Phyllophaga luctuosa	703324	species
Phyllophaga marginalis	1890715	species
- Phyllophaga micans	1890716	species
- Phyllophaga misteca	3273375	species
- Phyllophaga nebulosa	703325	species
- Phyllophaga nevermanni	1261310	species
- Phyllophaga nitida	1845068	species
- Phyllophaga nr. fraterna MP-2009	703331	species
- Phyllophaga orosina	1274977	species
- Phyllophaga panamana	1274978	species
- Phyllophaga parvidens	1890717	species
- Phyllophaga paternoi	703326	species
- Phyllophaga pearliae	703327	species
- Phyllophaga perlonga	1890718	species
- Phyllophaga pleei	903835	species
- Phyllophaga polyphylla	3248545	species
- Phyllophaga postrema	1890719	species
- Phyllophaga praetermissa	489810	species
- Phyllophaga profunda	489811	species
- Phyllophaga prolixa	1262764	species
- Phyllophaga pruinosa	1261311	species
- Phyllophaga prunina	489812	species
- Phyllophaga pseudofloridana	703328	species
- Phyllophaga ravida	3248546	species
- Phyllophaga reventazona	1261312	species
- Phyllophaga rubiginosa	489813	species
- Phyllophaga rugosa	1710757	species
- Phyllophaga schaefferi	1890720	species
- Phyllophaga schizorhina	1261313	species
- Phyllophaga setidorsis	1261314	species
- Phyllophaga sodalis	2680841	species
- Phyllophaga spreta	489814	species
- Phyllophaga submucida	489815	species
- Phyllophaga subtonsa	703329	species
- Phyllophaga tecta	703330	species
- Phyllophaga torta	489816	species
- Phyllophaga tristis	1710758	species
- Phyllophaga vetula	1225478	species
- Phyllophaga vilifrons	1890721	species
unclassified Phyllophaga	2634113	no rank
Phyllotocus	207283	genus
- Phyllotocus navicularis	365800	species
- Phyllotocus rufipennis	365687	species
unclassified Phyllotocus	2622601	no rank
Plectris	980945	genus
unclassified Plectris	2641722	no rank
Polyphylla	614209	genus
- Polyphylla adspersa	1074824	species
- Polyphylla albolineata	3095326	species
- Polyphylla barbata	692529	species
- Polyphylla decemlineata	1248204	species
- Polyphylla fullo	1476581	species
- Polyphylla gracilicornis	1618307	species
Polyphylla laticollis	1290871	species
- Polyphylla olivieri	614210	species
- Polyphylla schoenfeldti	3377368	species
- Polyphylla variolosa	2976700	species
- Polyphylla vietnamica	3087116	species
Prodontria	50565	genus
- Prodontria bicolorata	50572	species
- Prodontria capito	50571	species
- Prodontria lewisi	50570	species
- Prodontria longitarsis	50569	species
- Prodontria modesta	50568	species
- Prodontria pinguis	50573	species
- Prodontria praelatella	50574	species
- Prodontria truncata	50566	species
unclassified Prodontria	2642510	no rank
Pseudosymmachia	1618312	genus
- Pseudosymmachia flavescens	1618313	species
- Pseudosymmachia impressifrons	2859698	species
Pseudotrochalus	1695874	genus
- Pseudotrochalus quadrilineatus	1698185	species
unclassified Pseudotrochalus	2640902	no rank
Puelchesia	702051	genus
- Puelchesia gracilis	702052	species
Rhabdopholis	1237349	genus
unclassified Rhabdopholis	2636646	no rank
Rhizotrogus	1971887	genus
- Rhizotrogus aestivus	1971888	species
Rhopaea	272070	genus
- Rhopaea magnicornis	272071	species	brown cockchafer
unclassified Rhopaea	2635670	no rank
Scelophysa	1033764	genus
unclassified Scelophysa	2621324	no rank
Schizonycha	207289	genus
- Schizonycha abrupta	1237299	species
- Schizonycha affinis	1221088	species
- Schizonycha algoensis	1237318	species
- Schizonycha carbonaria	1237300	species
- Schizonycha cf. confinis DA-2012	1237241	species
- Schizonycha compacta	1237301	species
- Schizonycha debilis	1237303	species
- Schizonycha divulsa	1237304	species
- Schizonycha durbana	1237305	species
- Schizonycha fallax	1237319	species
- Schizonycha fuscescens	383528	species
- Schizonycha globa	1237306	species
- Schizonycha gracilis	1237307	species
- Schizonycha increta	1237308	species
- Schizonycha languens	1237309	species
- Schizonycha lebidis	1237310	species
- Schizonycha livada	1237311	species
- Schizonycha neglecta	1237312	species
- Schizonycha piceonigra	1237313	species
- Schizonycha puncticollis	1237314	species
- Schizonycha ruficollis	3374174	species
- Schizonycha transvaalica	1237315	species
- Schizonycha usambarae	1237316	species
- Schizonycha valida	1237317	species
unclassified Schizonycha	2629349	no rank
Scitala	1552087	genus
unclassified Scitala	2635850	no rank
Scitalia	365803	genus
- Scitalia aureorufa	365804	species
unclassified Scitalia	2999154	no rank
Scythrodes	50563	genus
- Scythrodes squalidus	50564	species
Selaserica	2945124	genus
- Selaserica athukoralai	2945156	species
- Selaserica convexiuscula	2945157	species
- Selaserica fabriziae	2945370	species
- Selaserica impexa	2945158	species
- Selaserica karnatakaensis	3402272	species
- Selaserica maculicauda	2945159	species
- Selaserica nitida	2945160	species
- Selaserica nuwarana	3069466	species
- Selaserica praetexta	3069467	species
- Selaserica pusilla	2945161	species
unclassified Selaserica	2945125	no rank
Sericesthis	1552092	genus
- Sericesthis nigrolineata	2681018	species
unclassified Sericesthis	2629028	no rank
Sericoides	478143	genus
unclassified Sericoides	2630846	no rank
Sophrops	383517	genus
- Sophrops heydeni	1822362	species
- Sophrops karschi	3379183	species
- Sophrops peronosporus	3064708	species
- Sophrops sculpticollis	3402273	species
- Sophrops striatus	761925	species
- Sophrops subrugatus	2962662	species
unclassified Sophrops	2639387	no rank
Sparrmannia	1208543	genus
unclassified Sparrmannia (in: beetles)	2688946	no rank
Sphaeroscelis	1552096	genus
unclassified Sphaeroscelis	2639710	no rank
Telura	1552098	genus
unclassified Telura	2633805	no rank
Teluroides	1552100	genus
unclassified Teluroides	2624197	no rank
Thyce	2681154	genus
- Thyce squamicollis	2681155	species
Toxospathius	3385920	genus
- Toxospathius auriventris	3385921	species
Trematodes	1290878	genus
- Trematodes tenebrioides	1290879	species
Trichomaladera	1695889	genus
unclassified Trichomaladera	2685798	no rank
Varencya	465355	genus
unclassified Varencya	2648068	no rank
Xylonichus	1552110	genus
unclassified Xylonichus	2635784	no rank
unclassified Melolonthinae	465027	no rank
- Enariini sp. MA DA-2012	1207341	species
- Hopliini sp. DA-2011	980935	species
- Madohopliini sp. BMNH 671507	465016	species
- Melolonthinae gen. 1 sp. 1 MAC-2021a	2904868	species
- Melolonthinae sp. BMNH 671352	465028	species
- Melolonthinae sp. BOLD:AAH0276	2659817	species
- Melolonthinae sp. BOLD:AAH0481	2659825	species
- Melolonthinae sp. BOLD:AAH0854	2659821	species
- Melolonthinae sp. BOLD:AAH0855	2659819	species
- Melolonthinae sp. BOLD:AAH0856	2659824	species
- Melolonthinae sp. BOLD:AAH0857	2659827	species
- Melolonthinae sp. BOLD:AAH0858	2659822	species
- Melolonthinae sp. BOLD:AAH0859	2659826	species
- Melolonthinae sp. BOLD:AAH0865	2659823	species
- Melolonthinae sp. BOLD:AAH0866	2659820	species
- Melolonthinae sp. BOLD:AAV2189	2659818	species
- Melolonthinae sp. BOLD:AAV2190	2659816	species
- Melolonthinae sp. BOLD:ACC1981	1710042	species
- Melolonthinae sp. NLG-2014	1551922	species
- Melolonthinae sp. c.s.syu201335	1822353	species
- Melolonthinae sp. c.s.syu201340	1822354	species
- Melolonthini sp. DA-2012	1207342	species
- Melolonthini sp. KE1 DA-2012	1207343	species
- Melolonthini sp. KE2 DA-2012	1207344	species
Orphninae	207266	subfamily
Hybalus	207267	genus
unclassified Hybalus	2639512	no rank
Madecorphnus	2729074	genus
- Madecorphnus grebennikovi	2729076	species
Orphnus	702312	genus
unclassified Orphnus	2621494	no rank
Paraegidium	2570520	genus
- Paraegidium costalimai	2570521	species
Rutelinae	7062	subfamily	shining leaf chafers
Adoretus	207257	genus
- Adoretus duvauceli	1475329	species
- Adoretus flavus	1475330	species
- Adoretus hirsutus	2795331	species
- Adoretus kanarensis	3049077	species
- Adoretus lasiopygus	383479	species
- Adoretus nasalis	3385660	species
- Adoretus ovalis	2871087	species
- Adoretus puberulus	1618291	species
- Adoretus sorex	3394782	species
- Adoretus tenuimaculatus	1078810	species
- Adoretus versutus	383484	species
unclassified Adoretus	2624234	no rank
Anisoplia	190490	genus
- Anisoplia austriaca	1962506	species
- Anisoplia depressa	190491	species
- Anisoplia dispar	465108	species
- Anisoplia lata	3144895	species
- Anisoplia remota	190492	species
- Anisoplia tempestiva	3182363	species
Anomala	71817	genus
- Anomala aereiventris	1940410	species
- Anomala aglaos	1940411	species
Anomala albopilosa	452287	species
- Anomala amphicoma	1261270	species
- Anomala antiqua	2055703	species
- Anomala arara	1940423	species
- Anomala aulacoides	1689724	species
- Anomala aulax	1198227	species
- Anomala balzapambae	1261271	species
- Anomala bengalensis	383485	species
- Anomala biharensis	383486	species
- Anomala bilobata	383487	species
- Anomala binotata	2680106	species
- Anomala calceuiformis	3388681	species
- Anomala calligrapha	1261272	species
- Anomala cantori	383488	species
- Anomala carlsoni	702041	species
- Anomala chamaeleon	761907	species
- Anomala chapini	1261273	species
- Anomala clarivillosa	1940412	species
- Anomala clathrata	1940424	species
- Anomala communis	2704554	species
- Anomala contusa	1940413	species
- Anomala corpulenta	931571	species
- Anomala costata	1069816	species
- Anomala cuprea	121601	species	cupreous chafer
- Anomala cuprea x Anomala albopilosa	3122384	species
- Anomala cupreovariolosa	1940425	species
- Anomala cupricollis	1261274	species
- Anomala dimidiata	383489	species
- Anomala discoidalis	1446079	species
- Anomala divisa	1940426	species
- Anomala dorsalis	383490	species
- Anomala dubia	1143021	species
- Anomala dussumieri	3126204	species
- Anomala eucoma	1261275	species
- Anomala eulissa	1940427	species
- Anomala eusticta	1940414	species
- Anomala exoleta	1618292	species
Anomala expansa	1453165	species
- Anomala ferrea	1940428	species
- Anomala flavacoma	1940429	species
- Anomala fuscosignata	3233837	species
- Anomala geniculata	3140909	species
- Anomala globulata	1940430	species
- Anomala grandis	1502669	species
- Anomala graueri	1745990	species
- Anomala hoppi	1940431	species
- Anomala innuba	2680107	species
- Anomala koreana	3140910	species
- Anomala levicollis	1940415	species
- Anomala lineatopennis	1475331	species
- Anomala longicarcarata	3388682	species
- Anomala longisacculata	1940416	species
- Anomala luculenta	761908	species
- Anomala luminosa	3233838	species
- Anomala marginata	2680108	species
- Anomala mausonica	3388683	species
- Anomala megalia	1446080	species
- Anomala megaparamera	1940432	species
- Anomala mesosticta	1940433	species
- Anomala mimeloides	1262762	species
- Anomala mongolica	761909	species
- Anomala moroni	1940434	species
- Anomala nigroflava	1940435	species
- Anomala obovata	1940436	species
- Anomala octiescostata	121602	species
- Anomala orientalis	1310277	species
- Anomala osakana	71818	species
- Anomala parvaeucoma	1940437	species
- Anomala perspicax	1940417	species
- Anomala pincelada	1940438	species
- Anomala polita	383491	species
- Anomala praecellens	1262763	species
- Anomala praenitens	383492	species
- Anomala pseudoeucoma	1940439	species
- Anomala quiche	1940440	species
Anomala rubripes	3233836	species
- Anomala ruficapilla	1075165	species
- Anomala rufiventris	3064707	species
- Anomala rufocuprea	121600	species	soybean beetle
- Anomala russiventris	2962660	species
- Anomala schoenfeldti	3095327	species
- Anomala schonfeldti	121603	species
- Anomala semicincta	1261276	species
- Anomala semilla	1940441	species
- Anomala sieversi	3140911	species
- Anomala singularis	3402268	species
- Anomala spatuliformis	3388684	species
- Anomala spiloptera	3233840	species
- Anomala stillaticia	1940418	species
- Anomala strigodermoides	1940442	species
- Anomala subridens	1940419	species
- Anomala tenoriensis	1940420	species
- Anomala testaceipennis	1940443	species
- Anomala trapezifera	1446081	species
- Anomala triformis	3233841	species
- Anomala tuberculata	1940421	species
- Anomala undulata	2680109	species
- Anomala valida	1446082	species
- Anomala vallisneria	1940422	species
- Anomala varians	2704555	species
- Anomala varicolor	383497	species
- Anomala variegata	383498	species
- Anomala viridana	2859683	species
- Anomala viridicostata	3233842	species
- Anomala vitalisi	3240851	species
- Anomala vitis	3230648	species
- Anomala volsellata	1940444	species
- Anomala vulcanicola	1282353	species
- Anomala xantholoma	3240852	species
- Anomala xanthoptera	383499	species
- Anomala zumbadoi	1940445	species
environmental samples	696123	no rank
unclassified Anomala	2611935	no rank
Anomalorhina	1446090	genus
- Anomalorhina turrialbana	1446091	species
Anoplognathus	302445	genus
- Anoplognathus boisduvalii	302446	species
- Anoplognathus olivieri	2680113	species
- Anoplognathus porosus	2680114	species
unclassified Anoplognathus	2622978	no rank
Anoplostethus	1551966	genus
unclassified Anoplostethus	2626201	no rank
Anticheira	980919	genus
unclassified Anticheira	2638743	no rank
Blithopertha	465185	genus
unclassified Blithopertha	2636531	no rank
Blitopertha	1630971	genus
- Blitopertha lineolata	2490418	species
Brachysternus	1298931	genus
- Brachysternus prasinus	1298932	species
Callistethus	1261328	genus
- Callistethus carbo	1447028	species
- Callistethus chontalensis	1261329	species
- Callistethus collaris	1261330	species
- Callistethus fuscorubens	1447029	species
- Callistethus granulipygus	1446083	species
- Callistethus lativittis	1447031	species
- Callistethus levigatus	1447032	species
- Callistethus macroxantholeus	1447033	species
- Callistethus marginicollis	1446089	species
- Callistethus microxantholeus	1447034	species
- Callistethus multiplicatus	1447035	species
- Callistethus parapulcher	1447036	species
- Callistethus plagiicollis	1618295	species
- Callistethus ruteloides	1940446	species
- Callistethus schneideri	1446084	species
- Callistethus specularis	1446085	species
- Callistethus stannibractea	1447037	species
- Callistethus valdecostatus	1447038	species
- Callistethus yalizo	1940447	species
unclassified Callistethus	2641886	no rank
Calloodes	1551982	genus
- Calloodes rayneri	2680196	species
unclassified Calloodes	2649215	no rank
Chaetopteroplia	1587442	genus
- Chaetopteroplia segetum	1587443	species
Chejuanomala	3140912	genus
- Chejuanomala quelparta	3140913	species
Chlorota	1261331	genus
- Chlorota panamensis	1261332	species
Chrysina	1261333	genus
- Chrysina aurigans	1261334	species
- Chrysina aurilisternum	1874195	species
- Chrysina aurora	1261335	species
- Chrysina bruyeai	1261336	species
- Chrysina gloriosa	3098083	species
- Chrysina limbata	1261337	species
- Chrysina luteomarginata	1261338	species
- Chrysina peruviana	1874196	species
- Chrysina resplendens	1897488	species
Cotalpa	1053876	genus
- Cotalpa lanigera	1053877	species
Exomala	121604	genus
- Exomala conspurcata	3402771	species
- Exomala orientalis	121605	species	oriental beetle
- Exomala pallidipennis	761899	species
Heteronychus	207264	genus
- Heteronychus arator	295550	species	African black beetle
- Heteronychus lioderes	383500	species
unclassified Heteronychus	2637667	no rank
Hylamorpha	1336370	genus
- Hylamorpha elegans	1336371	species
Isoplia	980907	genus
- Isoplia lasiosoma	980908	species
Kibakoganea	1631033	genus
- Kibakoganea sinica	3161192	species
- Kibakoganea tamdaoensis	3241133	species
Lepadoretus	465276	genus
- Lepadoretus ovatoides	465277	species
Mimadoretus	1552042	genus
unclassified Mimadoretus	2644797	no rank
Mimela	383515	genus
- Mimela holosericea	1618300	species
- Mimela junii	1588243	species
- Mimela siliguria	383524	species
- Mimela splendens	1618301	species
- Mimela testaceipes	1618302	species
- Mimela xanthorhina	3379182	species
unclassified Mimela	2621098	no rank
Oryctomorphus	1542261	genus
- Oryctomorphus bimaculatus	1542262	species
Paracotalpa	1631074	genus
- Paracotalpa puncticollis	2042168	species
- Paracotalpa ursina	2042169	species
Paraschizognathus	1609505	genus
unclassified Paraschizognathus	2638210	no rank
Parastasia	1247182	genus
Parastasia ferrieri	1247183	species
unclassified Parastasia	2612256	no rank
Pelidnota	575207	genus
- Pelidnota belti	1261297	species
- Pelidnota frommeri	1261298	species
- Pelidnota notata	1261299	species
- Pelidnota parallela	1261300	species
- Pelidnota punctata	575208	species
unclassified Pelidnota	2644921	no rank
Phalangogonia	1261352	genus
- Phalangogonia sperata	1261353	species
Phyllopertha	93415	genus
- Phyllopertha diversa	93416	species
- Phyllopertha horticola	207390	species
Platycoelia	1261354	genus
- Platycoelia humeralis	1274979	species
- Platycoelia mesosternalis	1261355	species
- Platycoelia puncticollis	2937924	species
Plusiotis	207285	genus
unclassified Plusiotis	2649956	no rank
Popillia	7063	genus
- Popillia flavosellata	1618308	species
- Popillia japonica	7064	species	Japanese beetle
- Popillia mutans	1453201	species
- Popillia quadriguttata	1618309	species
- Popillia taiwana	1453202	species
unclassified Popillia	2622461	no rank
Proagopertha	1630949	genus
- Proagopertha lucidula	1630950	species
Prodoretus	465335	genus
- Prodoretus truncatus	465336	species
Repsimus	1552082	genus
- Repsimus aeneus	2680980	species
unclassified Repsimus	2645379	no rank
Rutela	702053	genus
- Rutela sanguinolenta	702054	species
Schizognathus	1609507	genus
unclassified Schizognathus	2626466	no rank
Spodochlamys	1261360	genus
- Spodochlamys cupreola	1261361	species
Strigoderma	1262765	genus
- Strigoderma arbicola	1980619	species
- Strigoderma auriventris	1940449	species
unclassified Strigoderma	2641153	no rank
Trigonostomum	465342	genus
- Trigonostomum mucoreum	465343	species
Trioplognathus	1609509	genus
unclassified Trioplognathus	2618855	no rank
unclassified Rutelinae	465023	no rank
- Adoretini sp. BMNH 671419	465024	species
- Adoretini sp. BMNH 671443	1142395	species
- Rutelinae sp. BMNH 747070	465026	species
- Rutelinae sp. BOLD:AAG4511	2660822	species
Scarabaeidae incertae sedis	2617486	no rank
Calocolobopterus	2612860	genus
- Calocolobopterus principalis	2612861	species
Neocolobopterus	2613121	genus
- Neocolobopterus maculicollis	2613122	species
Trichaphodioides	2613293	genus
- Trichaphodioides pulchellus	2613294	species
unclassified Trichaphodioides	3382415	no rank
Scarabaeinae	41142	subfamily	dung beetles & tumblebugs
Ateuchini	569636	tribe
Ateuchus	206691	genus
Canthidium	206804	genus
Leotrichillum	1925861	genus
Trichillidium	1925800	genus
Australasian clade	2169428	clade
Amphistomus	206791	genus
Aptenocanthon	206573	genus
Canthonosoma	206815	genus
Cephalodesmius	206822	genus
Coproecus	206827	genus
Lepanus	206887	genus
Matthewsius	2169430	genus
Monteithocanthon	2169429	genus
Saphobius	206920	genus
Sauvagesinella	518541	genus
Tesserodon	518556	genus
Coprini	569635	tribe
Copris	41145	genus	black dung beetles
Coptodactyla	206833	genus
Homocopris	1925798	genus
Litocopris	1165017	genus
Deltochilini	569629	tribe
Aliuscanthoniola	1165013	genus
Anachalcos	206793	genus
Aphengoecus	518213	genus
Apotolamprus	369520	genus
Arachnodes	369525	genus
Bohepilissus	518221	genus
Boletoscapter	206802	genus
Byrrhidium	518226	genus
Cambefortantus	518429	genus
Canthon	206809	genus
Canthonella	518457	genus
Circellium	205290	genus
Cryptocanthon	518464	genus
Deltochilum	206838	genus
Diorygopyx	206855	genus
Dwesasilvasedis	763524	genus
Endroedyolus	518473	genus
Epilissus	490890	genus
Epirinus	206700	genus
Eudinopus	206862	genus
Gyronotus	518481	genus
Hammondantus	518482	genus
Hansreia	206877	genus
Haroldius	518483	genus
Ignambia	206885	genus
Labroma	518494	genus
Malagoniella	377170	genus
Megathopa	377172	genus
Megathoposoma	206891	genus
Monoplistes	206575	genus
Namakwanus	518506	genus
Nanos	490857	genus
Odontoloma	206703	genus
Oficanthon	518511	genus
Onthobium	206715	genus
Outenikwanus	518513	genus
Panelus	206717	genus
Paracanthon	518514	genus
Peckolus	518521	genus
Pseudignambia	206579	genus
Pseudocanthon	518532	genus
Pseudonthobium	206721	genus
Pycnopanelus	518539	genus
Saphobiamorpha	518540	genus
Scybalocanthon	206926	genus
Scybalophagus	206723	genus
Silvaphilus	1165019	genus
Tanzanolus	518555	genus
Temnoplectron	206928	genus
Tesserodoniella	518557	genus
Epactoidini	3021417	tribe
Epactoides	475445	genus
Grebennikovius	3073736	genus
Ochicanthon	475431	genus
Eucraniini	569630	tribe
Anomiopsoides	205288	genus
Ennearabdus	206860	genus
Eucranium	205294	genus
Glyphoderus	206873	genus
Eurysternini	569631	tribe
Eurysternus	206864	genus
Gymnopleurini	569632	tribe
Allogymnopleurus	206789	genus
Garreta	518479	genus
Gymnopleurus	166367	genus
Paragymnopleurus	206719	genus
Oniticellini	569639	tribe
Euoniticellus	166327	genus
Liatongus	206889	genus
Oniticellus	206900	genus
Tibiodrepanus	2795037	genus
Tiniocellus	206931	genus
Onthophagini	569640	tribe
Caccobius	166381	genus
Cleptocaccobius	206825	genus
Digitonthophagus	206852	genus
Euonthophagus	166364	genus
Eusaproecius	518475	genus
Hyalonthophagus	206883	genus
Milichus	206893	genus
Mimonthophagus	518505	genus
Onthophagus	166331	genus
Phalops	206911	genus
Proagoderus	206917	genus
Scarabaeinae incertae sedis	569643	no rank
Anomiopus	206796	genus
Anonthobium	206798	genus
Anonychonitis	518210	genus
Aptychonitis	518216	genus
Bdelyropsis	206800	genus
Bdelyrus	377167	genus
Bolbites	377168	genus
Bubas	166325	genus
Canthodimorpha	518440	genus
Catharsius	206817	genus
Cheironitis	166378	genus
Coprophanaeus	206694	genus
Coptorhina	206696	genus
Cyptochirus	518466	genus
Delopleurus	518467	genus
Demarziella	206844	genus
Dendropaemon	206848	genus
Diabroctis	206850	genus
Dichotomius	206698	genus
Dicranocara	392728	genus
Drepanocerus	206857	genus
Frankenbergerius	518478	genus
Gromphas	206875	genus
Helictopleurus	206879	genus
Heliocopris	205296	genus
Heteronitis	206881	genus
Janssensantus	518490	genus
Macroderes	206711	genus
Megatharsis	518502	genus
Metacatharsius	206713	genus
Microcopris	423537	genus
Neateuchus	205301	genus
Onitis	166329	genus
Ontherus	206903	genus
Orthophagus	3044103	genus
Oruscatus	377173	genus
Oxysternon	206905	genus
Pachysoma	205305	genus
Parachorius	518515	genus
Paracopris	422262	genus
Paraphytus	518518	genus
Parascatonomus	369558	genus
Paronthobium	206907	genus
Pedaria	206705	genus
Phanaeus	206913	genus
Pseudoarachnodes	369561	genus
Sarophorus	206923	genus
Scaptodera	1848583	genus
Scatimus	518543	genus
Serrophorus	422268	genus
Sulcophanaeus	377174	genus
Thyregis	518560	genus
Tragiscus	206934	genus
Trichillum	206725	genus
Uroxys	206707	genus
Xinidium	518566	genus
Scarabaeini	569633	tribe
Aulacium	2745772	genus
Drepanopodus	205292	genus
Kheper	205298	genus
Pachylomerus	205303	genus
Scarabaeus	41143	genus	scarabs
Sisyphini	569634	tribe
Neosisyphus	206895	genus
Nesosisyphus	518509	genus
Sisyphus	166371	genus
unclassified Scarabaeinae	518198	no rank
- Scarabaeinae gen. 1 sp. 1 NLG-2017	1970079	species
- Scarabaeinae gen. 2 sp. 2 NLG-2017	1970081	species
- Scarabaeinae sp.	3063634	species
- Scarabaeinae sp. BOLD:AAG4809	2660889	species
- Scarabaeinae sp. BOLD:AAH0281	2660888	species
- Scarabaeinae sp. BOLD:AAP6547	2660887	species
Sericinae	360072	subfamily
Ablaberoides	465164	genus
- Ablaberoides abyssinicus	1698064	species
- Ablaberoides cf. SA5 DA-2012	1237202	species
- Ablaberoides crassus	1698065	species
- Ablaberoides festiva	1698054	species
- Ablaberoides quasitus	1698055	species
- Ablaberoides tenellus	1698066	species
unclassified Ablaberoides	2624928	no rank
Allokotarsa	465166	genus
- Allokotarsa clypeata	465167	species
unclassified Allokotarsa	2647227	no rank
Amiserica	1631451	genus
- Amiserica antennalis	1698056	species
- Amiserica costulata	1698067	species
- Amiserica insperata	1698068	species
- Amiserica malickyi	1698070	species
- Amiserica rufidula	1698069	species
- Amiserica similissima	1698071	species
unclassified Amiserica	2632927	no rank
Anomalophylla	465169	genus
- Anomalophylla huashanica	1631501	species
- Anomalophylla tristicula	465170	species
unclassified Anomalophylla	2685403	no rank
Apotriodonta	614205	genus
- Apotriodonta hispanica	1698072	species
Archeohomaloplia	1698230	genus
- Archeohomaloplia ganhaiziensis	1698231	species
Arraphytarsa	1695840	genus
unclassified Arraphytarsa	2645171	no rank
Astaena	465176	genus
unclassified Astaena	2626689	no rank
Calloserica	1695897	genus
- Calloserica poggii	1698074	species
Chrysoserica	465201	genus
- Chrysoserica auricoma	1698075	species
- Chrysoserica stebnickae	465202	species
Comaserica	465205	genus
- Comaserica crinita	465206	species
unclassified Comaserica	2634613	no rank
Cyrtocamenta	465220	genus
unclassified Cyrtocamenta	2642609	no rank
Eriphoserica	1698232	genus
- Eriphoserica camentoides	1698233	species
Eumaladera	1631454	genus
- Eumaladera subrugata	1631478	species
unclassified Eumaladera	2635694	no rank
Euronycha	1237341	genus
- Euronycha cf. rhodesiana DA-2012	1237342	species
unclassified Euronycha	2646748	no rank
Euserica	465237	genus
- Euserica lucipeta	1698076	species
- Euserica mutata	1698077	species
- Euserica villarreali	1142406	species
Gastroserica	1631456	genus
- Gastroserica carolusi	2873613	species
- Gastroserica dembickyi	1698078	species
- Gastroserica fanjingensis	1631479	species
- Gastroserica guizhouana	1631480	species
- Gastroserica haucki	2873614	species
- Gastroserica herzi	1822360	species
- Gastroserica huaphanensis	2873615	species
- Gastroserica impressicollis	1631481	species
- Gastroserica kabakovi	1631482	species
- Gastroserica marginalis	1698079	species
- Gastroserica nigrofasciata	1698080	species
- Gastroserica viridis	1698081	species
unclassified Gastroserica	2623912	no rank
Glaphyserica	1698234	genus
- Glaphyserica humeralis	1698235	species
Gryphonycha	1698236	genus
- Gryphonycha puberula	1698237	species
Gynaecoserica	465245	genus
- Gynaecoserica aniniensis	1698082	species
- Gynaecoserica annuliforceps	1698083	species
- Gynaecoserica cymosa	1698084	species
- Gynaecoserica exilis	1698085	species
- Gynaecoserica hirsuta	1698086	species
- Gynaecoserica obliqua	1698087	species
- Gynaecoserica tawangensis	1698088	species
- Gynaecoserica variipennis	465246	species
unclassified Gynaecoserica	2634255	no rank
Hellaserica	465250	genus
- Hellaserica elongata	465251	species
Heteroserica	1698238	genus
- Heteroserica tridens	1698239	species
Hymenoplia	465258	genus
- Hymenoplia arragonica	681106	species
- Hymenoplia clypealis	681107	species
- Hymenoplia escalerai	681108	species
- Hymenoplia fulvipennis	478133	species
- Hymenoplia galaica	681109	species
- Hymenoplia lata	1698089	species
- Hymenoplia lineolata	465259	species
- Hymenoplia pseudocinerascens	681110	species
- Hymenoplia rugulosa	681111	species
unclassified Hymenoplia	2620615	no rank
Hyposerica	465261	genus
unclassified Hyposerica	2617896	no rank
Idaeserica	465263	genus
unclassified Idaeserica	2644315	no rank
Lamproserica	465272	genus
unclassified Lamproserica	2685415	no rank
Lasioserica	1631462	genus
- Lasioserica brevipilosa	1698090	species
- Lasioserica dragon	1631483	species
- Lasioserica kubani	1698091	species
- Lasioserica maculata	1698092	species
- Lasioserica meghalayana	2873616	species
- Lasioserica modikholae	1698093	species
- Lasioserica oblita	1698094	species
- Lasioserica pudens	1698095	species
unclassified Lasioserica	2627029	no rank
Lepiserica	465278	genus
- Lepiserica lydenburgiana	1704664	species
unclassified Lepiserica	2649729	no rank
Leuroserica	1698240	genus
- Leuroserica stemmleri	1698241	species
Maladera	360073	genus
- Maladera affinis	383504	species
- Maladera anderssoni	2945132	species
- Maladera annamensis	1698096	species
- Maladera atavana	1698097	species
- Maladera badullana	2945133	species
- Maladera bandarwelana	2945134	species
- Maladera basalis	465129	species
- Maladera bombycina	3394784	species
- Maladera breviatella	2945135	species
- Maladera burmeisteri	465130	species
- Maladera calcarata	2945136	species
- Maladera calicutensis	2745527	species
- Maladera cambodjana	1698098	species
- Maladera cardoni	383505	species
- Maladera cariniceps	3401916	species
- Maladera castanea	1254970	species
- Maladera cervicornis	2945137	species
- Maladera cf. affinis 835085	1698256	species
- Maladera cf. affinis 835100	1698255	species
- Maladera cf. fuscescens BMNH 671454	478134	species
- Maladera cf. fuscescens DA-2007	465131	species
- Maladera cf. fuscescens DA-2012	1237216	species
- Maladera cinnaberina	2945138	species
- Maladera clypeata	2873617	species
- Maladera coxalis	2945139	species
- Maladera dambullana	2945364	species
- Maladera drescheri	1698099	species
- Maladera dubia	2945140	species
- Maladera exima	2873618	species
- Maladera fistulosa	2945141	species
- Maladera formosae	2918028	species
- Maladera fumosa	3394785	species
- Maladera fuscescens	465132	species
- Maladera fuscipes	2873619	species
- Maladera futschauana	2873620	species
- Maladera galdaththana	2945142	species
- Maladera heveli	2945143	species
Maladera himalayica	465012	species
- Maladera hiyarensis	3069458	species
- Maladera hmong	1698100	species
- Maladera holosericea	360074	species
- Maladera hortonensis	2945144	species
- Maladera insanabilis	1475337	species
- Maladera insubida	1698101	species
- Maladera iridescens	465133	species
- Maladera iuga	2945365	species
- Maladera japonica	761920	species
- Maladera joachimi	465134	species
- Maladera kanarana	465135	species
- Maladera kandyensis	3069459	species
- Maladera karunaratnae	2945366	species
- Maladera kawaii	1698102	species
- Maladera kishi	2945367	species
- Maladera kreyenbergi	1698103	species
- Maladera lateritia	2945145	species
- Maladera lindulana	2945146	species
- Maladera lugubris	3394786	species
- Maladera magnicornis	3394787	species
- Maladera masumotoi	1698104	species
- Maladera miliouensis	1698105	species
- Maladera mollis	3069460	species
- Maladera murzini	1698106	species
- Maladera mutabilis	3394788	species
- Maladera obscurata	1698107	species
- Maladera okamotoi	761921	species
- Maladera orientalis	941252	species
- Maladera padaviyaensis	2945147	species
- Maladera peregoi	2873621	species
- Maladera perniciosa	1698108	species
- Maladera phoenicica	1698109	species
- Maladera praviforceps	3394789	species
- Maladera propagator	3394790	species
- Maladera pseudoexima	2873945	species
- Maladera pubescens	2945148	species
- Maladera quinquidens	383506	species
- Maladera rotunda	1631484	species
- Maladera rotundata	2945149	species
- Maladera rufocuprea	2945150	species
- Maladera rufodorsata	2873622	species
- Maladera rufoplagiata	2873623	species
- Maladera rufotestacea	1698110	species
- Maladera saitoi	1698111	species
- Maladera senta	2873624	species
- Maladera seriatoguttata	3394791	species
- Maladera setosa	2945151	species
- Maladera significans	465136	species
- Maladera simlana	465137	species
- Maladera siniaevi	1698112	species
- Maladera sontica	1698113	species
- Maladera spectabilis	1698114	species
- Maladera straminea	1698115	species
- Maladera stridula	1698116	species
- Maladera sujitrae	3402270	species
- Maladera taiwana	1631485	species
- Maladera taurica	1698117	species
- Maladera thirthahalliensis	3394792	species
- Maladera thomsoni	383507	species
- Maladera tibialis	1698118	species
- Maladera tricuspidata	2945152	species
- Maladera unguicularis	1698119	species
- Maladera ventriosa	3394793	species
- Maladera vernacula	3394794	species
- Maladera verticalis	1618286	species
- Maladera weligamana	2945153	species
- Maladera wewalkai	1698120	species
- Maladera windy	2945368	species
unclassified Maladera	2634803	no rank
Microserica	465008	genus
- Microserica affinis	1698121	species
- Microserica bhutanensis	1698122	species
- Microserica corporaali	1698123	species
- Microserica crenatostriata	1698124	species
- Microserica dohrni	1698125	species
- Microserica duplosetosa	2572776	species
- Microserica fukiensis	1631486	species
- Microserica gandakiensis	465145	species
- Microserica interrogator	465146	species
- Microserica iridicolor	1698126	species
- Microserica limbata	1705922	species
- Microserica lucens	1698127	species
- Microserica malaccensis	1698128	species
- Microserica panzona	2873625	species
- Microserica pruinosa	465009	species
- Microserica pulchella	1698129	species
- Microserica pyrrhopoecila	1698130	species
- Microserica simplex	2873626	species
- Microserica singalangia	1698131	species
- Microserica soppongensis	465147	species
- Microserica varians	1698132	species
- Microserica viridicollis	1698133	species
- Microserica vittipennis	1698134	species
unclassified Microserica	2638925	no rank
Microtrochalus	1698242	genus
- Microtrochalus plagiger	1698243	species
Neomaladera	1698244	genus
- Neomaladera barbara	1698245	species
Neoserica	465293	genus
- Neoserica allolaotica	1698135	species
- Neoserica arunachalana	1698057	species
- Neoserica aulica	3402271	species
- Neoserica baishuiensis	1698136	species
- Neoserica barbara	1698138	species
- Neoserica barberi	2750230	species
- Neoserica biuncinata	1698139	species
- Neoserica cardamomensis	1698140	species
- Neoserica carneola	1698141	species
- Neoserica dharmapriyai	2945154	species
- Neoserica flavoviridis	3394795	species
- Neoserica fukiensis	1631487	species
- Neoserica gravida	3394796	species
- Neoserica guangpingensis	1698144	species
- Neoserica lateriuncinata	1698149	species
- Neoserica lucidifrons	1698058	species
- Neoserica martinui	2873627	species
- Neoserica mukdahan	1698152	species
- Neoserica ningyuanensis	1698153	species
- Neoserica phuphanensis	2873628	species
- Neoserica pophami	2945369	species
- Neoserica probsti	1698156	species
- Neoserica pubiforceps	1698158	species
- Neoserica rubiginea	3092634	species
- Neoserica ruzickai	1698160	species
- Neoserica septemlamellata	1698161	species
- Neoserica sexfoliata	2945155	species
- Neoserica silvestris	1698164	species
- Neoserica speciosa	1698165	species
- Neoserica tamdaoensis	1698059	species
- Neoserica transvaalica	1698166	species
- Neoserica ursina	1631488	species
- Neoserica vicina	1698060	species
unclassified Neoserica	2622356	no rank
Nepaloserica	465304	genus
- Nepaloserica hartmanni	1698172	species
- Nepaloserica mustangia	465305	species
- Nepaloserica pewaensis	1698173	species
- Nepaloserica procera	1698174	species
- Nepaloserica rufescens	1698175	species
- Nepaloserica schmidti	465306	species
unclassified Nepaloserica	2619917	no rank
Neuroserica	465311	genus
- Neuroserica cf. fulvescens/mashona DA-2012	1237225	species
- Neuroserica fulvescens	465312	species
- Neuroserica mashona	465313	species
Nipponoserica	478147	genus
- Nipponoserica koltzei	478148	species
- Nipponoserica peregrina	3059121	species
unclassified Nipponoserica	2647161	no rank
Omaloplia	360076	genus
- Omaloplia nigromarginata	465148	species
- Omaloplia ruricola	465149	species
Oxyserica	465318	genus
- Oxyserica pygidialis	465319	species
Pachyserica	465322	genus
- Pachyserica albosignata	1631489	species
- Pachyserica albosquamosa	1698176	species
- Pachyserica janbezdeki	1698177	species
- Pachyserica minax	3231386	species
- Pachyserica olafi	465323	species
unclassified Pachyserica	2626441	no rank
Paramaladera	1698246	genus
- Paramaladera major	1698247	species
- Paramaladera rufofusca	1698248	species
Paratriodonta	478145	genus
- Paratriodonta cinctipennis	1698178	species
- Paratriodonta lentula	1698179	species
- Paratriodonta normandi	1698180	species
- Paratriodonta romana	478146	species
unclassified Paratriodonta	2620158	no rank
Pleophylla	1205641	genus
- Pleophylla burundiensis	3114071	species
- Pleophylla charlyi	3114072	species
- Pleophylla congoensis	3114073	species
- Pleophylla fasciatipennis	1698181	species
- Pleophylla ferruginea	1698182	species
- Pleophylla harrisoni	1816084	species
- Pleophylla kruegeri	3114074	species
- Pleophylla lizleri	3114075	species
- Pleophylla mlilwaneensis	3114076	species
- Pleophylla mpumalanga	3114077	species
- Pleophylla murzini	3114078	species
- Pleophylla navicularis	1698183	species
- Pleophylla nelshoogteensis	1816085	species
- Pleophylla pilosa	1698184	species
- Pleophylla pseudopilosa	1816086	species
- Pleophylla ruthae	3114079	species
- Pleophylla settentrionalis	3114080	species
- Pleophylla silvatica	1816087	species
- Pleophylla tongaatsana	3114081	species
- Pleophylla transkeiensis	3114082	species
- Pleophylla wakkerstroomensis	2792801	species
- Pleophylla warnockae	1816088	species
unclassified Pleophylla	2630589	no rank
Serica	360075	genus
- Serica acontia	1698186	species
- Serica alleni	1698187	species
- Serica alternata	1698188	species
- Serica atracapilla	1710790	species
- Serica atrata	465153	species
- Serica benesi	465154	species
- Serica brunnea	465155	species
- Serica campestris	2681014	species
- Serica concinna	2681015	species
- Serica curvata	1710791	species
- Serica daliensis	1631490	species
- Serica eberti	1698189	species
- Serica elmontea	1698190	species
- Serica emeishanica	1631491	species
- Serica erectosetosa	1698191	species
- Serica falcata	1698192	species
- Serica fimbriata	1698193	species
- Serica formosana	1705918	species
- Serica furcata	1698194	species
- Serica fusa	2945162	species
- Serica fusifemorata	1698195	species
Serica georgiana	2083700	species
- Serica heydeni	465156	species
- Serica intermixta	465158	species
- Serica jindrai	1631492	species
- Serica karnaliensis	1698196	species
- Serica khajiaris	1698197	species
- Serica kingdoni	1631493	species
- Serica lagoi	1698198	species
- Serica ligulata	1698199	species
- Serica lijiangensis	1698200	species
- Serica loxia	465159	species
- Serica lurida	2945163	species
- Serica maculosa	1631494	species
- Serica mystica	465160	species
- Serica nebulosa	1631495	species
- Serica nigroguttata	1698201	species
- Serica ochrosoma	2681016	species
- Serica panchaseana	1698202	species
- Serica parallela	1710792	species
- Serica perigonia	1698203	species
- Serica porcula	2681017	species
- Serica sculptilis	1698204	species
- Serica sericea	1710793	species
- Serica solivaga	1631496	species
- Serica sponsa	2486051	species
- Serica stygia	1698205	species
- Serica sudhausi	1631497	species
- Serica thibetana	1698206	species
- Serica tristis	1483839	species
- Serica tryznai	1631498	species
- Serica tukucheana	465163	species
- Serica werneri	2873629	species
- Serica wrasei	1631499	species
Taiwanoserica	1695887	subgenus
unclassified Serica	2623739	no rank
Sericania	1695885	genus
- Sericania fuscolineata	2673498	species
unclassified Sericania	2637215	no rank
Tetraserica	465337	genus
- Tetraserica appendiculata	2570850	species
- Tetraserica auriculata	2570841	species
- Tetraserica bachmaensis	2570866	species
- Tetraserica chiangdaoensis	2570848	species
- Tetraserica curviforceps	2570844	species
- Tetraserica damaidiensis	3092636	species
- Tetraserica doisangensis	2570854	species
- Tetraserica falciformis	2570843	species
- Tetraserica ferrugata	1698209	species
- Tetraserica filiforceps	2570845	species
- Tetraserica kontumensis	2570864	species
- Tetraserica leishanica	2962663	species
- Tetraserica liangheensis	1698210	species
- Tetraserica loeiensis	2570859	species
- Tetraserica longipenis	2570847	species
- Tetraserica maerimensis	2570861	species
- Tetraserica matsumotoi	2570856	species
- Tetraserica namnaoensis	2570858	species
- Tetraserica neouncinata	2570846	species
- Tetraserica pahinngamensis	2570853	species
- Tetraserica pailinensis	2570840	species
- Tetraserica petrpacholatkoi	2570865	species
- Tetraserica pingjiangensis	1698061	species
- Tetraserica pseudoruiliensis	2570851	species
- Tetraserica pseudouncinata	2570842	species
- Tetraserica romae	2570862	species
- Tetraserica shangsiensis	3092635	species
- Tetraserica spinicrus	2570849	species
- Tetraserica takahashii	2570863	species
- Tetraserica tanahrataensis	2570852	species
- Tetraserica wapiensis	2570855	species
- Tetraserica wiangpapaoana	2570860	species
- Tetraserica yaoquensis	2572777	species
unclassified Tetraserica	2635226	no rank
Triodontella	465344	genus
Triodontella alni	1698211	species
- Triodontella lajonquierei	1698213	species
- Triodontella mimula	1698214	species
- Triodontella nitidula	1698215	species
- Triodontella raymondi	1698216	species
- Triodontella zuzartei	1698217	species
unclassified Triodontella	2618743	no rank
Trioserica	1631467	genus
- Trioserica maculipennis	1631500	species
unclassified Trioserica	2621083	no rank
Trochaloserica	465351	genus
- Trochaloserica festina	478142	species
unclassified Trochaloserica	2631774	no rank
Trochalus	465353	genus
- Trochalus aeruginosus	1698062	species
- Trochalus diversirostris	1698218	species
- Trochalus exasperans	1698219	species
- Trochalus ferrugineus	1698220	species
- Trochalus lepidus	1698221	species
- Trochalus splendidulus	1698063	species
- Trochalus vagus	1698222	species
unclassified Trochalus	2622200	no rank
Xenoserica	1698249	genus
- Xenoserica brachyptera	1698250	species
unclassified Sericinae	465010	no rank
- Sericinae sp. SAX DA-2012	1243157	species
- Trochalina sp. BMNH 671448	465025	species
- cf. Microserica sp. DA-2007	465011	species
Trichiinae	174902	subfamily
Gnorimus	347352	genus
- Gnorimus decempunctatus	1207086	species
- Gnorimus nobilis	1207087	species
- Gnorimus subopacus	2673474	species
- Gnorimus variabilis	347353	species
Valginae	207364	subfamily
Valgus	207365	genus
- Valgus canaliculatus	1542198	species
- Valgus hemipterus	207366	species
unclassified Scarabaeidae	260547	no rank
- Scarabaeidae sp.	2807411	species
- Scarabaeidae sp. 8104_004	2545402	species
- Scarabaeidae sp. BAH_28362_D08	2588018	species
- Scarabaeidae sp. BAH_28362_G09	2588122	species
- Scarabaeidae sp. BAH_28362_G10	2588019	species
- Scarabaeidae sp. BAH_28362_H06	2588121	species
- Scarabaeidae sp. BIOUG02561-C02	1989977	species
- Scarabaeidae sp. BIOUG05376-H10	1998998	species
- Scarabaeidae sp. BIOUG17834-H05	1989978	species
- Scarabaeidae sp. BIOUG18623-E03	1995994	species
- Scarabaeidae sp. BIOUG18623-E08	1998997	species
- Scarabaeidae sp. BM677796	397659	species
- Scarabaeidae sp. BM677798	397660	species
- Scarabaeidae sp. BMNH 1274750	1796539	species
- Scarabaeidae sp. BMNH 1274752	1796540	species
- Scarabaeidae sp. BMNH 1274753	1796541	species
- Scarabaeidae sp. BOLD:AAC0491	2791844	species
- Scarabaeidae sp. BOLD:AAF0418	1845915	species
- Scarabaeidae sp. BOLD:AAG9045	2660886	species
- Scarabaeidae sp. BOLD:AAH0159	1845916	species
- Scarabaeidae sp. BOLD:AAH6793	1845917	species
- Scarabaeidae sp. BOLD:AAH6798	1845918	species
- Scarabaeidae sp. BOLD:AAH6827	1845919	species
- Scarabaeidae sp. BOLD:AAM7660	1845920	species
- Scarabaeidae sp. BOLD:AAM7691	2660878	species
- Scarabaeidae sp. BOLD:AAP6956	2660876	species
- Scarabaeidae sp. BOLD:AAP6957	2660879	species
- Scarabaeidae sp. BOLD:AAP7124	2791845	species
- Scarabaeidae sp. BOLD:AAP7127	2791846	species
- Scarabaeidae sp. BOLD:AAP7128	2791847	species
- Scarabaeidae sp. BOLD:AAP7129	2791848	species
- Scarabaeidae sp. BOLD:AAP7130	2791849	species
- Scarabaeidae sp. BOLD:AAP7131	2791850	species
- Scarabaeidae sp. BOLD:AAP7134	2791851	species
- Scarabaeidae sp. BOLD:AAP7135	2791852	species
- Scarabaeidae sp. BOLD:AAP7136	2791853	species
- Scarabaeidae sp. BOLD:AAP7139	2791854	species
- Scarabaeidae sp. BOLD:AAP7140	2791855	species
- Scarabaeidae sp. BOLD:AAP7142	2791856	species
- Scarabaeidae sp. BOLD:AAP7143	2791857	species
- Scarabaeidae sp. BOLD:AAP7144	2791858	species
- Scarabaeidae sp. BOLD:AAP7878	2791859	species
- Scarabaeidae sp. BOLD:AAP7879	2791860	species
- Scarabaeidae sp. BOLD:AAP7881	2791862	species
- Scarabaeidae sp. BOLD:AAP7882	2791863	species
- Scarabaeidae sp. BOLD:AAP7883	2791864	species
- Scarabaeidae sp. BOLD:AAP7888	2791865	species
- Scarabaeidae sp. BOLD:AAP7895	2791866	species
- Scarabaeidae sp. BOLD:AAP7896	2791867	species
- Scarabaeidae sp. BOLD:AAP7897	2791868	species
- Scarabaeidae sp. BOLD:AAP7911	2791869	species
- Scarabaeidae sp. BOLD:AAP7914	2791870	species
- Scarabaeidae sp. BOLD:AAP7915	2791871	species
- Scarabaeidae sp. BOLD:AAP7916	2791872	species
- Scarabaeidae sp. BOLD:AAP7917	2791873	species
- Scarabaeidae sp. BOLD:AAP7918	2791874	species
- Scarabaeidae sp. BOLD:AAP7919	2791875	species
- Scarabaeidae sp. BOLD:AAP7920	2791876	species
- Scarabaeidae sp. BOLD:AAP7921	2791877	species
- Scarabaeidae sp. BOLD:AAP7933	2791878	species
- Scarabaeidae sp. BOLD:AAP7934	2791879	species
- Scarabaeidae sp. BOLD:AAP7935	2791880	species
- Scarabaeidae sp. BOLD:AAP7936	2791881	species
- Scarabaeidae sp. BOLD:AAP7946	2791882	species
- Scarabaeidae sp. BOLD:AAP7949	2791883	species
- Scarabaeidae sp. BOLD:AAP7952	2791884	species
- Scarabaeidae sp. BOLD:AAP7953	2791885	species
- Scarabaeidae sp. BOLD:AAP7954	2791886	species
- Scarabaeidae sp. BOLD:AAP7959	2791887	species
- Scarabaeidae sp. BOLD:AAP7960	2791888	species
- Scarabaeidae sp. BOLD:AAP7961	2791889	species
- Scarabaeidae sp. BOLD:AAP7962	2791890	species
- Scarabaeidae sp. BOLD:AAP7966	2791891	species
- Scarabaeidae sp. BOLD:AAP7971	2791892	species
- Scarabaeidae sp. BOLD:AAP7981	2791893	species
- Scarabaeidae sp. BOLD:AAP7991	2791894	species
- Scarabaeidae sp. BOLD:AAP8590	2791895	species
- Scarabaeidae sp. BOLD:AAP8591	2791896	species
- Scarabaeidae sp. BOLD:AAP8592	2791897	species
- Scarabaeidae sp. BOLD:AAP8593	2791898	species
- Scarabaeidae sp. BOLD:AAP8599	2791899	species
- Scarabaeidae sp. BOLD:AAP8612	2791900	species
- Scarabaeidae sp. BOLD:AAP8792	2791901	species
- Scarabaeidae sp. BOLD:AAP8793	2791902	species
- Scarabaeidae sp. BOLD:AAP8794	2791903	species
- Scarabaeidae sp. BOLD:AAP8795	2791904	species
- Scarabaeidae sp. BOLD:AAP9990	2660885	species
- Scarabaeidae sp. BOLD:AAP9991	2660874	species
- Scarabaeidae sp. BOLD:AAP9992	2660881	species
- Scarabaeidae sp. BOLD:AAP9993	2660884	species
- Scarabaeidae sp. BOLD:AAP9994	2660883	species
- Scarabaeidae sp. BOLD:AAP9996	2660875	species
- Scarabaeidae sp. BOLD:AAP9997	2660873	species
- Scarabaeidae sp. BOLD:AAP9998	2660877	species
- Scarabaeidae sp. BOLD:AAQ3307	2660880	species
- Scarabaeidae sp. BOLD:AAY9690	1845921	species
- Scarabaeidae sp. BOLD:ABA9089	1710152	species
- Scarabaeidae sp. BOLD:ABY7886	2792278	species
- Scarabaeidae sp. BOLD:ACE7729	2660882	species
- Scarabaeidae sp. CCDB-30443-F07	2612341	species
- Scarabaeidae sp. CCDB-30443-H06	2612340	species
- Scarabaeidae sp. CNC COL EO119834	2917908	species
- Scarabaeidae sp. DNAS-1DC-211949	2175048	species
- Scarabaeidae sp. DNAS-297-224277	2267352	species
- Scarabaeidae sp. DSP-2011a	1254578	species
- Scarabaeidae sp. DSP-2011b	1254579	species
- Scarabaeidae sp. INDOBIOSYS-CCDB24596-D01	2505497	species
- Scarabaeidae sp. INDOBIOSYS-CCDB24596-D04	2505498	species
- Scarabaeidae sp. INDOBIOSYS-CCDB24596-D11	2505499	species
- Scarabaeidae sp. INDOBIOSYS-CCDB24683-B09	2505496	species
- Scarabaeidae sp. INDOBIOSYS-CCDB25261-H11	2505495	species
- Scarabaeidae sp. INDOBIOSYS-CCDB25321-A07	2515293	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26008-C06	2505504	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26008-E11	2505500	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26008-F12	2505502	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26008-G06	2505503	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26209-A09	2505501	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26209-D07	2505494	species
- Scarabaeidae sp. INDOBIOSYS-CCDB26585-A02	2515292	species
- Scarabaeidae sp. NS_04	2866973	species
- Scarabaeidae sp. NZAC 03006214	1597691	species
- Scarabaeidae sp. NZAC 03006283	1597692	species
- Scarabaeidae sp. NZAC 03009063	1597693	species
- Scarabaeidae sp. NZAC 03009140	1597694	species
- Scarabaeidae sp. NZAC 03009233	1788218	species
- Scarabaeidae sp. NZAC 03009294	1597695	species
- Scarabaeidae sp. NZAC 03010439	1597696	species
- Scarabaeidae sp. NZAC 03010484	1597697	species
- Scarabaeidae sp. NZAC 03010514	1597698	species
- Scarabaeidae sp. NZAC 03010650	1597699	species
- Scarabaeidae sp. cvb777prey013	2559363	species
- Scarabaeidae sp. sc_05783	1901769	species
Trogidae	110048	family
Omorgus	295553	genus
- Omorgus alternans	1461450	species
- Omorgus asperulatus	463069	species
- Omorgus borrei	1461451	species
- Omorgus candidus	463070	species
- Omorgus cf. asperulatus YvdM-2007	463071	species
- Omorgus chinensis	2841931	species
- Omorgus costatus	463072	species
- Omorgus demarzi	463073	species
- Omorgus freyi	1461446	species
- Omorgus gemmatus	1461447	species
- Omorgus melancholicus	1461448	species
- Omorgus monachus	1461452	species
- Omorgus pastillarius	1462748	species
- Omorgus pauliani	1461449	species
- Omorgus punctatus	2680755	species
- Omorgus radula	463074	species
- Omorgus spatulatus	1461453	species
- Omorgus squalidus	463076	species
- Omorgus squamosus	295554	species
- Omorgus subcarinatus	1461454	species
- Omorgus suberosus	463077	species
- Omorgus tatei	1461455	species
- Omorgus tesselatus	1464068	species
- Omorgus trilobus	1461456	species
unclassified Omorgus	2630513	no rank
- Omorgus sp. ACS7130	2892711	species
- Omorgus sp. BMNH 793032	1142414	species
- Omorgus sp. CSR057	2055474	species
- Omorgus sp. NLG-2014	1551928	species
- Omorgus sp. YvdM-2007	463075	species
Phoberus	1846235	genus
- Phoberus aculeatus	1660358	species
- Phoberus arcuatus	463081	species
- Phoberus brincki	1461458	species
Phoberus caffer	463082	species
- Phoberus caffer caffer	463083	subspecies
- Phoberus capensis	463084	species
- Phoberus consimilis	1461460	species
- Phoberus cyrtus	761657	species
- Phoberus disjunctus	1846236	species
Phoberus fascicularis	463085	species
- Phoberus fascicularis fascicularis	1461472	subspecies
- Phoberus fascicularis rowei	463086	subspecies
- Phoberus herminae	1846237	species
- Phoberus horridus	1461462	species
- Phoberus luridus	1461463	species
- Phoberus montanus	463089	species
- Phoberus mozalae	1461473	species
- Phoberus nama	761658	species
- Phoberus nanniscus	463090	species
- Phoberus nasutus	1461465	species
- Phoberus natalensis	1461466	species
- Phoberus planicollis	463093	species
- Phoberus pusillus	1660359	species
- Phoberus quadricostatus	1461474	species
- Phoberus rhyparoides	463094	species
- Phoberus rudebecki	295552	species
- Phoberus squamiger	463098	species
- Phoberus strigosus	1461470	species
- Phoberus sulcatus	463099	species
- Phoberus talpa	463100	species
- Phoberus youngai	1660360	species
Polynoncus	463059	genus
- Polynoncus brevicollis	463078	species
- Polynoncus bullatus	463079	species
Trox	186085	genus
- Trox aequalis	463080	species
- Trox atrox	2917892	species
- Trox brahminus	1461457	species
Trox cadaverinus	1461459	species
- Trox cadaverinus cadaverinus	1461471	subspecies
- Trox fabricii	1461461	species
- Trox gemmulatus	463087	species
- Trox hamatus	463088	species
- Trox hispidus	1971873	species
- Trox mandli	1461464	species
- Trox niponensis	463091	species
- Trox opacotuberculatus	463092	species
- Trox perlatus	1461467	species
- Trox perrieri	1461468	species
- Trox perrisii	1586555	species
- Trox sabulosus	1553806	species
- Trox scaber	1461469	species
Trox setifer	463095	species
- Trox setifer setifer	463096	subspecies
- Trox spinulosis	463097	species
- Trox terrestris	463101	species
- Trox tuberculatus	463102	species
- Trox uenoi	273925	species
- Trox unistriatus	219498	species
- Trox ussuriensis	1461475	species
- Trox variolatus	1896867	species
unclassified Trox	2641199	no rank
- Trox sp. AD-2015	1661510	species
- Trox sp. BMNH 678430	465216	species
- Trox sp. BMNH 793026	1142425	species
- Trox sp. BMNH 793028	1142426	species
- Trox sp. BMNH 793031	1142427	species
- Trox sp. C41	186086	species
- Trox sp. CRS 2021	2848625	species
- Trox sp. INB036	2055564	species
- Trox sp. JH-2005	358432	species
- Trox sp. TRO01	1205588	species
unclassified Trogidae	1597752	no rank
- Trogidae sp.	2900080	species
- Trogidae sp. BOLD:ACJ0539	1846111	species
- Trogidae sp. NZAC03011909	1788284	species
- Trogidae sp. NZAC03011913	1788285	species
unclassified Scarabaeoidea	702044	no rank
- Scarabaeoidea sp. KM-2017	2219446	species